黔中高原喀斯特常绿与落叶阔叶混交林类型及群落特征
贺忠权, 刘长成, 蔡先立, 郭柯

Types and community characteristics of karst mixed evergreen and deciduous broad-leaved forests in the central Guizhou Plateau, China
HE Zhong-Quan, LIU Chang-Cheng, CAI Xian-Li, GUO Ke
表1 黔中高原喀斯特常绿与落叶阔叶混交林植被类型
Table 1 Vegetation types of karst mixed evergreen and deciduous broad-leaved forests in the central Guizhou Plateau
群系组 Alliance Group 群系 Alliance 群丛 Association 凭证样方
Plot number
青冈、落叶阔叶混交林Cyclobalanopsis and Deciduous broadleaf forest Alliance Group 青冈+圆果化香树林
Cyclobalanopsis glauca + Platycarya longipes Forest Alliance
青冈+圆果化香树-针齿铁仔-求米草群丛
Cyclobalanopsis glauca + Platycarya longipes - Myrsine semiserrata - Oplismenus undulatifolius Association
PD-S19
青冈+圆果化香树-针齿铁仔-江南星蕨群丛
Cyclobalanopsis glauca + Platycarya longipes - Myrsine semiserrata - Neolepisorus fortunei
Association
GZ2019-3
青冈+圆果化香树-狭叶海桐-求米草群丛
Cyclobalanopsis glauca + Platycarya longipes - Pittosporum glabratum var. neriifolium - Oplismenus undulatifolius Association
GZ2019-4
青冈+圆果化香树-青冈-西南莩草群丛
Cyclobalanopsis glauca + Platycarya longipes - Cyclobalanopsis glauca - Setaria forbesiana Association
GZ2013-33, GZ2013-34
青冈+云贵鹅耳枥林
Cyclobalanopsis glauca +
Carpinus pubescens
Forest
Alliance
青冈+云贵鹅耳枥-针齿铁仔-求米草群丛
Cyclobalanopsis glauca + Carpinus pubescens - Myrsine semiserrata - Oplismenus undulatifolius Association
PD-S7
青冈+云贵鹅耳枥-针齿铁仔-阔叶山麦冬群丛
Cyclobalanopsis glauca + Carpinus pubescens - Myrsine semiserrata - Liriope muscari Association
PD-S58
青冈+云贵鹅耳枥-云贵鹅耳枥-大披针薹草群丛
Cyclobalanopsis glauca + Carpinus pubescens - Carpinus pubescens - Carex lanceolata Association
QZ2019-5
青冈+云贵鹅耳枥-南天竹-江南星蕨群丛
Cyclobalanopsis glauca + Carpinus pubescens - Nandina domestica - Neolepisorus fortunei Association
QZ2019-8
青冈+朴树林
Cyclobalanopsis glauca +
Celtis sinensis
Forest Alliance
青冈+朴树-针齿铁仔-江南星蕨群丛
Cyclobalanopsis glauca + Celtis sinensis - Myrsine semiserrata -
Neolepisorus fortunei
Association
PD-S17
青冈+榉树林
Cyclobalanopsis glauca + Zelkova
serrata
Forest Alliance
青冈+榉树-香叶树-蝴蝶花群丛
Cyclobalanopsis glauca + Zelkova serrata - Lindera communis - Iris japonica Association
GZ2019-7
细叶青冈+圆果化香树林
Cyclobalanopsis gracilis +
Platycarya longipes Forest
Alliance
细叶青冈+圆果化香树-罗浮槭-大披针薹草群丛
Cyclobalanopsis gracilis + Platycarya longipes - Acer fabri - Carex lanceolata Association
QZ2019-1
细叶青冈+圆果化香树-针齿铁仔-阔叶山麦冬群丛
Cyclobalanopsis gracilis + Platycarya longipes - Myrsine semiserrata - Liriope muscari Association
QZ2019-2
群系组 Alliance Group 群系 Alliance 群丛 Association 凭证样方
Plot number
细叶青冈+圆果化香树-针齿铁仔-求米草群丛
Cyclobalanopsis gracilis + Platycarya longipes - Myrsine semiserrata - Oplismenus undulatifolius Association
PD-S18
细叶青冈+圆果化香树-针齿铁仔-对马耳蕨群丛
Cyclobalanopsis gracilis + Platycarya longipes - Myrsine semiserrata - Polystichum tsus-simense Association
PD-S20
细叶青冈+圆果化香树-细叶青冈-冷水花群丛
Cyclobalanopsis gracilis + Platycarya longipes - Cyclobalanopsis gracilis - Pilea notata Association
PD-S62
鼠刺、落叶阔叶混交林Itea and Deciduous broadleaf forest
Alliance Group
滇鼠刺+圆果化香树林
Itea yunnanensis + Platycarya longipes Forest Alliance
滇鼠刺+圆果化香树-圆果化香树-大披针薹草群丛
Itea yunnanensis + Platycarya longipes - Platycarya longipes -
Carex lanceolata
Association
GZ2007-2, GZ2013-8
滇鼠刺+圆果化香树-圆果化香树-荩草群丛
Itea yunnanensis + Platycarya longipes - Platycarya longipes -
Arthraxon hispidus
Association
GZ2007-3
滇鼠刺+圆果化香树-铁仔-大披针薹草群丛
Itea yunnanensis + Platycarya longipes - Myrsine africana -
Carex lanceolata
Association
GZ2009-2
滇鼠刺+圆果化香树-窄叶柯-求米草群丛
Itea yunnanensis + Platycarya longipes - Lithocarpus confinis -
Oplismenus undulatifolius
Association
GZ2009-7
滇鼠刺+圆果化香树-圆果化香树-求米草群丛
Itea yunnanensis + Platycarya longipes - Platycarya longipes -
Oplismenus undulatifolius
Association
GZ2009-14
滇鼠刺+圆果化香树-倒卵叶旌节花-顶芽狗脊群丛
Itea yunnanensis + Platycarya longipes - Stachyurus obovatus -
Woodwardia unigemmata
Association
GZ2013-9, PD-S60
滇鼠刺+圆果化香树-刺异叶花椒-大披针薹草群丛
Itea yunnanensis + Platycarya longipes - Zanthoxylum dimorphophyllum var. spinifolium - Carex lanceolata Association
PD-S24
滇鼠刺+圆果化香树-窄叶柯-大披针薹草群丛
Itea yunnanensis + Platycarya longipes - Lithocarpus confinis - Carex lanceolata Association
PD-S104
滇鼠刺+云贵鹅耳枥林
Itea yunnanensis + Carpinus
pubescens
Forest Alliance
滇鼠刺+云贵鹅耳枥-针齿铁仔-相近石韦群丛
Itea yunnanensis + Carpinus pubescens - Myrsine semiserrata -
Pyrrosia assimilis
Association
PD-S16
滇鼠刺+云贵鹅耳枥-青冈-西南莩草群丛
Itea yunnanensis + Carpinus pubescens - Cyclobalanopsis glauca -
Setaria forbesiana
Association
GZ2013-49
滇鼠刺+云贵鹅耳枥-山蜡梅-三枝九叶草群丛
Itea yunnanensis + Carpinus pubescens - Chimonanthus nitens -
Epimedium sagittatum
Association
GZ2013-57
滇鼠刺+云贵鹅耳枥-油茶-柄果薹草群丛
Itea yunnanensis + Carpinus pubescens - Camellia oleifera - Carex
stipitinux
Association
GZ2013-59
滇鼠刺+多脉猫乳林
Itea yunnanensis + Rhamnella
martini
Forest Alliance
滇鼠刺+多脉猫乳-刺异叶花椒-鸢尾群丛
Itea yunnanensis + Rhamnella martini - Zanthoxylum dimorphophyllum var. spinifolium - Iris tectorum Association
GZ2013-56
滇鼠刺+多脉猫乳-小冻绿树-大披针薹草群丛
Itea yunnanensis + Rhamnella martini - Rhamnus rosthornii - Carex lanceolata Association
PD-S56
滇鼠刺+栓皮栎林
Itea yunnanensis + Quercus
variabilis
Forest Alliance
滇鼠刺+栓皮栎-黑弹树-求米草群丛
Itea yunnanensis + Quercus variabilis - Celtis bungeana - Oplismenus undulatifolius Association
GZ2013-11
滇鼠刺+苦枥木林
Itea yunnanensis + Fraxinus
insularis
Forest Alliance
滇鼠刺+苦枥木-青冈-西南莩草群丛
Itea yunnanensis + Fraxinus insularis - Cyclobalanopsis glauca - Setaria forbesiana Association
GZ2013-48
樟、落叶阔叶混交林
Cinnamomum and
Deciduous broadleaf forest Alliance Group
猴樟+白栎林
Cinnamomum bodinieri + Quercus fabri Forest Alliance
猴樟+白栎-油茶-大披针薹草群丛
Cinnamomum bodinieri + Quercus fabri - Camellia oleifera - Carex lanceolata Association
PD-S11
群系组 Alliance Group 群系 Alliance 群丛 Association 凭证样方
Plot number
猴樟+白栎-朴树-大披针薹草群丛
Cinnamomum bodinieri + Quercus fabri - Celtis sinensis - Carex
lanceolata
Association
PD-S29
猴樟+白栎-香叶树-大披针薹草群丛
Cinnamomum bodinieri + Quercus fabri - Lindera communis - Carex lanceolata Association
PD-S64
猴樟+白栎-刺楸-大披针薹草群丛
Cinnamomum bodinieri + Quercus fabri - Kalopanax septemlobus - Carex lanceolata Association
PD-S85
猴樟+白栎-小冻绿树-庐山石韦群丛
Cinnamomum bodinieri + Quercus fabri - Rhamnus rosthornii - Pyrrosia sheareri Association
PD-S102
猴樟+圆果化香树林
Cinnamomum bodinieri + Platycarya longipes Forest Alliance
猴樟+圆果化香树-异叶梁王茶-顶芽狗脊群丛
Cinnamomum bodinieri + Platycarya longipes - Metapanax davidii - Woodwardia unigemmata Association
GZ2013-18
猴樟+圆果化香树-短萼海桐-丝叶薹草群丛
Cinnamomum bodinieri + Platycarya longipes - Pittosporum brevicalyx - Carex capilliformis Association
GZ2013-17
猴樟+响叶杨林
Cinnamomum bodinieri + Populus adenopoda Forest Alliance
猴樟+响叶杨-慈竹-顶芽狗脊群丛
Cinnamomum bodinieri + Populus adenopoda - Bambusa emeiensis - Woodwardia unigemmata Association
PD-S103
猴樟+栓皮栎林
Cinnamomum bodinieri + Quercus variabilis Forest Alliance
猴樟+栓皮栎-薄叶鼠李-狗脊群丛
Cinnamomum bodinieri + Quercus variabilis - Rhamnus leptophylla - Woodwardia unigemmata Association
PD-S66
猴樟+麻栎林
Cinnamomum bodinieri + Quercus acutissima Forest Alliance
猴樟+麻栎-朴树-渐尖毛蕨群丛
Cinnamomum bodinieri + Quercus acutissima - Celtis sinensis -
Cyclosorus acuminatus
Association
PD-S69
猴樟+椿叶花椒林
Cinnamomum bodinieri + Zanthoxylum ailanthoides Forest Alliance
猴樟+椿叶花椒-云贵鹅耳枥-大披针薹草群丛
Cinnamomum bodinieri + Zanthoxylum ailanthoides - Carpinus pubescens - Carex lanceolata Association
PD-S99
猴樟+翅荚香槐林
Cinnamomum bodinieri + Cladrastis platycarpa Forest Alliance
猴樟+翅荚香槐-香叶树-中华薹草群丛
Cinnamomum bodinieri + Cladrastis platycarpa - Lindera communis - Carex chinensis Association
GZ2013-13
柯、落叶阔叶混交林Lithocarpus and
Deciduous broadleaf forest Alliance Group
窄叶柯+圆果化香树林
Lithocarpus confinis + Platycarya longipes Forest Alliance
窄叶柯+圆果化香树-针齿铁仔-相近石韦群丛
Lithocarpus confinis + Platycarya longipes - Myrsine semiserrata -
Pyrrosia assimilis
Association
PD-S21
窄叶柯+圆果化香树-针齿铁仔-皱叶狗尾草群丛
Lithocarpus confinis + Platycarya longipes - Myrsine semiserrata -
Setaria plicata
Association
GZ2019-2
窄叶柯+圆果化香树-窄叶柯-薹草群丛
Lithocarpus confinis + Platycarya longipes - Lithocarpus confinis - Carex sp. Association
PDKZH-01
窄叶柯+圆果化香树-窄叶柯-丝叶薹草群丛
Lithocarpus confinis + Platycarya longipes - Lithocarpus confinis - Carex capilliformis Association
GZ2013-10
窄叶柯+圆果化香树-圆果化香树-千里光群丛
Lithocarpus confinis + Platycarya longipes - Platycarya longipes -
Senecio scandens
Association
GZ2009-12
窄叶柯+圆果化香树-香叶树-薹草群丛
Lithocarpus confinis + Platycarya longipes - Lindera communis - Carex sp. Association
QZ2019-7
窄叶柯+云贵鹅耳枥林
Lithocarpus confinis + Carpinus
pubescens
Forest Alliance
窄叶柯+云贵鹅耳枥-窄叶柯-大披针薹草群丛
Lithocarpus confinis + Carpinus pubescens - Lithocarpus confinis - Carex lanceolata Association
GZ2010-3
窄叶柯+云贵鹅耳枥-铁仔-中华薹草群丛
Lithocarpus confinis + Carpinus pubescens - Myrsine africana - Carex chinensis Association
GZ2013-6
窄叶柯+翅荚香槐林
Lithocarpus confinis + Cladrastis platycarpa Forest Alliance
窄叶柯+翅荚香槐-窄叶柯-薹草群丛
Lithocarpus confinis + Cladrastis platycarpa - Lithocarpus confinis - Carex sp. Association
QZ2019-3
群系组 Alliance Group 群系 Alliance 群丛 Association 凭证样方
Plot number
窄叶柯+翅荚香槐-川钓樟-江南星蕨群丛
Lithocarpus confinis + Cladrastis platycarpa - Lindera pulcherrima var. hemsleyana - Neolepisorus fortunei Association
PD-S4
润楠、落叶阔叶混交林Machilus and Deciduous broadleaf forest
Alliance Group
小果润楠+云贵鹅耳枥林
Machilus microcarpa + Carpinus
pubescens
Forest Alliance
小果润楠+云贵鹅耳枥-针齿铁仔-舌叶薹草群丛
Machilus microcarpa + Carpinus pubescens - Myrsine semiserrata - Carex ligulata Association
PD-S13
小果润楠+云贵鹅耳枥-川钓樟-大披针薹草群丛
Machilus microcarpa + Carpinus pubescens - Lindera pulcherrima var. hemsleyana - Carex lanceolata Association
PD-S8
小果润楠+云贵鹅耳枥-川钓樟-蝴蝶花群丛
Machilus microcarpa + Carpinus pubescens - Lindera pulcherrima var. hemsleyana - Iris japonica Association
QZ2019-6
小果润楠+圆果化香树林
Machilus microcarpa + Platycarya longipes Forest Alliance
小果润楠+圆果化香树-大果卫矛-江南星蕨群丛
Machilus microcarpa + Platycarya longipes - Euonymus myrianthus - Neolepisoru fortunei Association
PD-S88
小果润楠+圆果化香树-刺异叶花椒-渐尖毛蕨群丛
Machilus microcarpa + Platycarya longipes - Zanthoxylum
dimorphophyllum
var. spinifolium - Cyclosorus acuminatus Association
PD-S22
小果润楠+栓皮栎林
Machilus microcarpa + Quercus
variabilis
Forest Alliance
小果润楠+栓皮栎-窄叶柯-西南莩草群丛
Machilus microcarpa + Quercus variabilis - Lithocarpus confinis -
Setaria forbesiana
Association
GZ2013-12
宜昌润楠+圆果化香树林
Machilus ichangensis + Platycarya longipes Forest Alliance
宜昌润楠+圆果化香树-宜昌润楠-求米草群丛
Machilus ichangensis + Platycarya longipes - Machilus ichangensis - Oplismenus undulatifolius Association
GZ2013-36
安顺润楠+翅荚香槐林
Machilus cavaleriei + Cladrastis
platycarpa
Forest Alliance
安顺润楠+翅荚香槐-安顺润楠-求米草群丛
Machilus cavaleriei + Cladrastis platycarpa - Machilus cavaleriei - Oplismenus undulatifolius Association
GZ2013-14
石楠、落叶阔叶混交林Photinia and Deciduous broadleaf forest
Alliance Group
贵州石楠+朴树林
Photinia bodinieri +
Celtis sinensis
Forest Alliance
贵州石楠+朴树-朴树-阔叶山麦冬群丛
Photinia bodinieri + Celtis sinensis - Celtis sinensis - Liriope muscari Association
GZ2013-53
贵州石楠+朴树-香叶树-狭叶鳞毛蕨群丛
Photinia bodinieri + Celtis sinensis - Lindera communis - Dryopteris angustifrons Association
PD-S12
贵州石楠+梓林
Photinia bodinieri +
Catalpa ovata
Forest Alliance
贵州石楠+梓-野扇花-鸢尾群丛
Photinia bodinieri + Catalpa ovata - Sarcococca ruscifolia - Iris
tectorum
Association
GZ2013-50
贵州石楠+皂荚林
Photinia bodinieri + Gleditsia sinensis Forest Alliance
贵州石楠+皂荚-野扇花-鸢尾群丛
Photinia bodinieri + Gleditsia sinensis - Sarcococca ruscifolia -
Iris tectorum
Association
GZ2013-51
贵州石楠+刺楸林
Photinia bodinieri + Kalopanax
septemlobus
Forest Alliance
贵州石楠+刺楸-刺楸-求米草群丛
Photinia bodinieri + Kalopanax septemlobus - Kalopanax septemlobus - Oplismenus undulatifolius Association
GZ2019-1
贵州石楠+翅荚香槐林
Photinia bodinieri + Cladrastis
platycarpa
Forest Alliance
贵州石楠+翅荚香槐-香叶树-土人参群丛
Photinia bodinieri + Cladrastis platycarpa - Lindera communis -
Talinum paniculatum
Association
GZ2013-54
光叶石楠+圆果化香树林
Photinia glabra + Platycarya longipes Forest Alliance
光叶石楠+圆果化香树-针齿铁仔-抱石莲群丛
Photinia glabra + Platycarya longipes - Myrsine semiserrata -
Lemmaphyllum drymoglossoides
Association
GZ2013-55
山胡椒、落叶阔叶混交林 Lindera and Deciduous broadleaf forest Alliance Group 香叶树+山槐林
Lindera communis + Albizia kalkora Forest Alliance
香叶树+山槐-香叶树-麦冬群丛
Lindera communis + Albizia kalkora - Lindera communis - Ophiopogon japonicus Association
PD-S10
香叶树+响叶杨林
Lindera communis + Populus
adenopoda
Forest Alliance
香叶树+响叶杨-香叶树-顶芽狗脊群丛
Lindera communis + Populus adenopoda - Lindera communis -
Woodwardia unigemmata
Association
PD-S34
香叶树+麻栎林
Lindera communis + Quercus
acutissima
Forest Alliance
香叶树+麻栎-香叶树-丝叶薹草群丛
Lindera communis + Quercus acutissima - Lindera communis - Carex
capilliformis
Association
GZ2013-35
黑壳楠+梾木林
Lindera megaphylla + Cornus
macrophylla
Forest Alliance
黑壳楠+梾木-针齿铁仔-中华复叶耳蕨群丛
Lindera megaphylla + Cornus macrophylla - Myrsine semiserrata -
Arachniodes chinensis
Association
PD-S35
川钓樟+翅荚香槐林
Lindera pulcherrima var. hemsleyana + Cladrastis platycarpa Forest Alliance
川钓樟+翅荚香槐-川钓樟-大披针薹草群丛
Lindera pulcherrima var. hemsleyana + Cladrastis platycarpa - Lindera
pulcherrima
var. hemsleyana - Carex lanceolata Association
PD-S55
群系组 Alliance Group 群系 Alliance 群丛 Association 凭证样方
Plot number
海桐、落叶阔叶混交林Pittosporum and
Deciduous broadleaf forest Alliance Group
短萼海桐+翅荚香槐林
Pittosporum brevicalyx + Cladrastis platycarpa Forest Alliance
短萼海桐+翅荚香槐-小冻绿树-麦冬群丛
Pittosporum brevicalyx + Cladrastis platycarpa - Rhamnus rosthornii -
Ophiopogon japonicus
Association
PD-S71
短萼海桐+翅荚香槐-朴树-日本蛇根草群丛
Pittosporum brevicalyx + Cladrastis platycarpa - Celtis sinensis -
Ophiorrhiza japonica
Association
PD-S38
短萼海桐+圆果化香树林
Pittosporum brevicalyx + Platycarya longipes Forest Alliance
短萼海桐+圆果化香树-圆果化香树-求米草群丛
Pittosporum brevicalyx + Platycarya longipes - Platycarya longipes -
Oplismenus undulatifolius
Association
GZ2009-1
短萼海桐+栓皮栎林
Pittosporum brevicalyx + Quercus variabilis Forest Alliance
短萼海桐+栓皮栎-针齿铁仔-江南星蕨群丛
Pittosporum brevicalyx + Quercus variabilis - Myrsine semiserrata -
Neolepisorus fortunei
Association
PD-S61
槭、落叶阔叶混交林
Acer and Deciduous broadleaf forest
Alliance Group
罗浮槭+云贵鹅耳枥林
Acer fabri + Carpinus pubescens
Forest Alliance
罗浮槭+云贵鹅耳枥-异叶梁王茶-石筋草群丛
Acer fabri + Carpinus pubescens - Metapanax davidii - Pilea
plataniflora
Association
GZ2019-8
罗浮槭+云贵鹅耳枥-罗浮槭-大披针薹草群丛
Acer fabri + Carpinus pubescens - Acer fabri - Carex lanceolata Association
PD-S2
罗浮槭+圆果化香树林
Acer fabri + Platycarya longipes Forest Alliance
罗浮槭+圆果化香树-罗浮槭-蝴蝶花群丛
Acer fabri + Platycarya longipes - Acer fabri - Iris japonica Association
GZ2019-12
常绿栎、落叶
阔叶混交林
Quercus and Deciduous broadleaf forest
Alliance Group
乌冈栎+圆果化香树林
Quercus phillyreoides + Platycarya longipes Forest Alliance
乌冈栎+圆果化香树-乌冈栎-庐山石韦群丛
Quercus phillyreoides + Platycarya longipes - Quercus phillyreoides -
Pyrrosia sheareri
Association
GZ2019-6
巴东栎+白蜡树林
Quercus engleriana + Fraxinus
chinensis
Forest Alliance
巴东栎+白蜡树-巴东栎-中华复叶耳蕨群丛
Quercus engleriana + Fraxinus chinensis - Quercus engleriana -
Arachniodes chinensis
Association
QZ2019-4
杜英、落叶阔叶混交林Elaeocarpus and
Deciduous broadleaf forest Alliance Group
日本杜英+圆果化香树林
Elaeocarpus japonicus + Platycarya longipes Forest Alliance
日本杜英+圆果化香树-针齿铁仔-十字薹草群丛
Elaeocarpus japonicus + Platycarya longipes - Myrsine semiserrata -
Carex cruciata
Association
GZ2019-11