色季拉山不同腐解等级华山松倒木上的苔藓植物组合

doi:10.17521/cjpe.2019.0327

植物生态学报 ›› 2020, Vol. 44 ›› Issue (8): 842-853.DOI: 10.17521/cjpe.2019.0327 cstr: 32100.14.cjpe.2019.0327

所属专题：有机农业；苔藓生态

色季拉山不同腐解等级华山松倒木上的苔藓植物组合

刘凌¹, 樊英杰¹, 宋晓彤¹, 李敏², 邵小明¹, 王晓蕊³^,^*()

¹中国农业大学资源与环境学院, 生物多样性与有机农业北京市重点实验室, 北京 100193
²河北师范大学生命科学学院, 石家庄 050024
³石家庄学院资源与环境科学学院, 石家庄 050035

收稿日期:2019-11-25 接受日期:2020-04-27 出版日期:2020-08-20 发布日期:2020-06-12
作者简介:* wxr_wxr@126.com
基金资助:
国家自然科学基金(41771054);国家自然科学基金(31570474);河北省自然科学基金(D2019106006)

Bryophyte societies on the fallen logs of Pinus armandii with different decay classes in Sygera Mountains

LIU Ling¹, FAN Ying-Jie¹, SONG Xiao-Tong¹, LI Min², SHAO Xiao-Ming¹, WANG Xiao-Rui³^,^*()

¹College of Resources and Environmental Sciences, Beijing Key Laboratory of Biodiversity and Organic Farming, China Agricultural University, Beijing 100193, China
²College of Life Sciences, Hebei Normal University, Shijiazhuang 050024, China
³College of Resources and Environmental Science, Shijiazhuang University, Shijiazhuang 050035, China

Received:2019-11-25 Accepted:2020-04-27 Online:2020-08-20 Published:2020-06-12
Supported by:
National Natural Science Foundation of China(41771054);National Natural Science Foundation of China(31570474);Natural Science Foundation of Hebei Province(D2019106006)

摘要/Abstract

摘要：

倒木是森林生态系统维持健康和更新的重要组成部分, 在倒木不同腐解阶段, 倒木上定植的苔藓植物组合的差异尚不清楚。为深入探讨倒木上苔藓植物组合变化规律及其影响因素, 该文对西藏色季拉山同一地点不同腐解程度的4株华山松(Pinus armandii)倒木上的苔藓植物进行了样方调查和数据分析。结果表明: 4株倒木上40个样方共有苔藓植物22科52种, 其中藓类13科38种, 苔类9科14种; 以恒有优势种为特征进行划分, 共得到14个物种组合。随着倒木腐解程度加大以及周围环境的不断改变, 生长其上的苔藓植物物种多样性逐渐增加, 但组合数量和苔藓总盖度却呈现先增大后减小的趋势。苔藓物种由耐旱性强的丛生型藓类, 如木灵藓科、白发藓科, 逐渐演替为喜阴湿环境、快速生长的垫状或毯状藓类, 如提灯藓科、羽藓科和青藓科。倒木苔藓植物组合空间结构复杂程度逐渐增大。倒木是多种苔藓植物选择的重要生长基质, 其上苔藓植物受倒木的腐解程度、水分和光照等条件的影响而发生相应的物种组合变化。

关键词: 苔藓植物, 倒木, 生活型, 物种多样性, 环境因子

Abstract:

Aims Fallen logs play important roles in maintaining the health and the renewal of forest ecosystems, but the difference of bryophyte societies on fallen logs with different decay classes is unclear.
Methods To understand the regularity and influencing factors of changes in bryophyte societies, we investigated the bryophytes with quadrats along the trunks on four fallen logs of Pinus armandii with different decay classes at the same site in Sygera Mountains, Xizang. Then, the measured data were collected and analyzed.
Important findings There were 22 families and 52 species of bryophytes inhabiting the four fallen logs, including 13 families and 38 species of Musci, 9 families and 14 species of Hepaticae. According to the characteristics of the shared dominant species, 14 societies were obtained. With the increase of decay classes of fallen logs and changes in surrounding environment, the bryophyte diversity increased gradually, while the number of societies and the total bryophyte coverage fluctuated. Bryophyte species varied from drought-tolerant tall turfs, such as Orthotrichaceae and Leucobryaceae, to fast-growing cushions or mats in humid and shady environments, such as Mniaceae, Thuidiaceae and Brachytheciaceae. The complexity of spatial structure of bryophytic society increased gradually. Fallen logs are important growth substrates for a variety of bryophytes. Bryophyte societies on fallen logs vary with decay classes, moisture and light conditions.

Key words: bryophyte, fallen log, life form, biodiversity, environmental factor

刘凌, 樊英杰, 宋晓彤, 李敏, 邵小明, 王晓蕊. 色季拉山不同腐解等级华山松倒木上的苔藓植物组合. 植物生态学报, 2020, 44(8): 842-853. DOI: 10.17521/cjpe.2019.0327

LIU Ling, FAN Ying-Jie, SONG Xiao-Tong, LI Min, SHAO Xiao-Ming, WANG Xiao-Rui. Bryophyte societies on the fallen logs of Pinus armandii with different decay classes in Sygera Mountains. Chinese Journal of Plant Ecology, 2020, 44(8): 842-853. DOI: 10.17521/cjpe.2019.0327

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链接本文: https://www.plant-ecology.com/CN/10.17521/cjpe.2019.0327

https://www.plant-ecology.com/CN/Y2020/V44/I8/842

图/表 6

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倒木 Fallen log	倒木1 Log 1	倒木2 Log 2	倒木3 Log 3	倒木4 Log 4
针叶 Needle	存在 Present	无 Absent	无 Absent	无 Absent
树皮 Bark	与活体无显著区别 No difference from living	少部分脱落 Partial shedding	仅在粗大树枝上残存 Partly present only on large branches	无 Absent
树枝 Branches and twig	小树枝均存在 Twigs present	中等树枝存在 Branches present	粗大树枝存在 Large branches present	仅粗大树枝基部存在 Only large branches stub present
主干形状 Bole shape	圆形 Round	圆形 Round	圆形 Round	圆形至卵形 Round to oval
与地面位置 Position of log on ground	被某点抬高 Elevated on support point	被某点抬高 Elevated on support point	接近地面 Near or on the ground	整体落在地面 All of log on the ground
结构完整性 Structural integrity	木质体新鲜, 刀片可刺进 1-2 mm Cambium still fresh, knife blade penetrates 1-2 mm	开始腐解, 刀片可刺进 3 mm到1 cm Cambium decayed, knife blade penetrates 3 mm to 1 cm	刀片可刺进约2 cm Knife blade penetrates less than 2 cm	严重腐烂, 刀片可刺进 2-5 cm Severely decay, knife penetrates 2-5 cm
腐解等级 Decay class	Ⅰ	Ⅱ	Ⅲ	Ⅳ
胸径 Diameter at breast height	35 cm	23 cm	32 cm	27 cm
倒伏方向 Loging direction	南偏东15° 15° south by east	南偏东25° 25° south by east	南偏东15° 15° south by east	南偏东10° 10° south by east
倒木位置 Position of log	林窗中心 Center of forest gap	林窗中心 Center of forest gap	林内 Under the forest	林窗边缘^Edge of forest gap^
遮阴程度 Shade degree	10%	50%	90%	60%^*

倒木 Fallen log	倒木1 Log 1	倒木2 Log 2	倒木3 Log 3	倒木4 Log 4
针叶 Needle	存在 Present	无 Absent	无 Absent	无 Absent
树皮 Bark	与活体无显著区别 No difference from living	少部分脱落 Partial shedding	仅在粗大树枝上残存 Partly present only on large branches	无 Absent
树枝 Branches and twig	小树枝均存在 Twigs present	中等树枝存在 Branches present	粗大树枝存在 Large branches present	仅粗大树枝基部存在 Only large branches stub present
主干形状 Bole shape	圆形 Round	圆形 Round	圆形 Round	圆形至卵形 Round to oval
与地面位置 Position of log on ground	被某点抬高 Elevated on support point	被某点抬高 Elevated on support point	接近地面 Near or on the ground	整体落在地面 All of log on the ground
结构完整性 Structural integrity	木质体新鲜, 刀片可刺进 1-2 mm Cambium still fresh, knife blade penetrates 1-2 mm	开始腐解, 刀片可刺进 3 mm到1 cm Cambium decayed, knife blade penetrates 3 mm to 1 cm	刀片可刺进约2 cm Knife blade penetrates less than 2 cm	严重腐烂, 刀片可刺进 2-5 cm Severely decay, knife penetrates 2-5 cm
腐解等级 Decay class	Ⅰ	Ⅱ	Ⅲ	Ⅳ
胸径 Diameter at breast height	35 cm	23 cm	32 cm	27 cm
倒伏方向 Loging direction	南偏东15° 15° south by east	南偏东25° 25° south by east	南偏东15° 15° south by east	南偏东10° 10° south by east
倒木位置 Position of log	林窗中心 Center of forest gap	林窗中心 Center of forest gap	林内 Under the forest	林窗边缘^Edge of forest gap^
遮阴程度 Shade degree	10%	50%	90%	60%^*

科 Family	物种 Species	代码 Code	生活型 Life form	相对盖度 Relative coverage (%)
科 Family	物种 Species	代码 Code	生活型 Life form	Log 1	Log 2	Log 3	Log 4
曲背藓科 Oncophoraceae	曲背藓 Oncophorus wahlenbergii	OW	T-turf	9.74	-	-	-
曲背藓科 Oncophoraceae	南亚合睫藓 Symblepharis reinwardtii	SR	T-turf	-	-	-	1.99
	合睫藓 Symblepharis vaginata	SV	T-turf	-	44.18	-	2.99
曲尾藓科 Dicranaceae	波叶曲尾藓 Dicranum polysetum	DP	T-turf	-	-	3.97	-
白发藓科 Leucobryaceae	丛叶青毛藓 Dicranodontium caespitosum	DC	T-turf	-	8.52	4.77	6.97
	青毛藓 Dicranodontium denudatum	DiDe	T-turf	-	6.39	-	36.82
	长叶青毛藓 Dicranodontium didymodon	DiDi	T-turf	-	3.69	0.64	5.47
丛藓科 Pottiaceae	高山红叶藓 Bryoerythrophyllum alpigenum	BA	S-turf	-	-	0.32	-
	齿叶薄齿藓 Leptodontium handelii	LH	S-turf	0.49	-	-	-
	狭叶拟合睫藓 Pseudosymblepharis angustata	PA	S-turf	-	0.14	4.45	4.23
真藓科 Bryaceae	拟大叶真藓 Bryum salakense	BryS	T-turf	-	-	0.32	-
提灯藓科 Mniaceae	侧枝匐灯藓 Plagiomnium maximoviczii	PM	L-cushion	-	-	8.10	-
	全缘匐灯藓 Plagiomnium integrum	PI	L-cushion	-	-	-	1.00
	钝叶匐灯藓 Plagiomnium rostratum	PR	L-cushion	-	-	7.15	-
	具缘提灯藓 Mnium marginatum	MM	L-cushion	-	-	2.54	-
木灵藓科 Orthotrichaceae	毛帽木灵藓 Orthotrichum dasymitrium	OD	D-form	13.92	-	-	-
白齿藓科 Leucodontaceae	中华白齿藓 Leucodon sinensis	LS	Weft	11.68	-	-	-
蔓藓科 Meteoriaceae	新丝藓 Neodicladiella pendula	NP	R-mat	11.68	-	-	-
羽藓科 Thuidiaceae	毛羽藓 Bryonoguchia molkenboeri	BM	R-mat	-	-	-	0.50
羽藓科 Thuidiaceae	大羽藓 Thuidium cymbifolium	TC	R-mat	-	12.07	34.95	9.70
	毛尖羽藓 Thuidium plumulosum	TP	R-mat	-	-	-	0.50
	狭叶小羽藓 Haplocladium angustifolium	HA	R-mat	-	0.28	-	-
	细叶小羽藓 Haplocladium microphyllum	HM	R-mat	-	-	8.26	5.22
青藓科 Brachytheciaceae	多褶青藓 Brachythecium buchananii	BB	L-cushion	-	0.21	6.04	0.50
青藓科 Brachytheciaceae	平枝青藓 Brachythecium helminthocladum	BH	L-cushion	-	-	2.38	-
	燕尾青藓 Brachythecium novae-angliae	BraN	L-cushion	-	-	4.29	-
	褶叶青藓 Brachythecium salebrosum	BraS	L-cushion	-	-	-	1.24
	拟异叶藓 Pseudokindbergia dumosa	PD	L-cushion	-	-	0.64	-
灰藓科 Hypnaceae	圆枝粗枝藓 Gollania tereticaulis	GT	R-mat	-	2.13	-	-
灰藓科 Hypnaceae	灰藓 Hypnum cupressiforme	HC	R-mat	-	-	2.22	0.50
	密枝灰藓 Hypnum densirameum	HD	R-mat	-	2.13	2.86	7.46
	毛梳藓 Ptilium crista-castrensis	PC	R-mat	14.61	-	-	-
	拟硬叶藓 Stereodontopsis pseudorevoluta	SP	R-mat	0.19	-	-	-
	丝金灰藓 Pylaisia levieri	PL	R-mat	34.08	7.39	-	-
毛锦藓科 Pylaisiadelphaceae	曲叶小锦藓 Brotherella curvirostris	BC	R-mat	-	2.70	1.59	2.24
毛锦藓科 Pylaisiadelphaceae	弯叶小锦藓 Brotherella falcata	BF	R-mat	0.97	-	-	-
	垂蒴小锦藓 Brotherella nictans	BroN	R-mat	-	-	2.38	7.46
	暗绿毛锦藓 Pylaisiadelpha tristoviridis	PT	R-mat	-	4.12	-	-
叶苔科 Jungermanniaceae	深绿叶苔 Jungermannia atrovirens	JA	D-form	-	4.69	0.16	1.24
叶苔科 Jungermanniaceae	小萼叶苔 Jungermannia parviperiantha	JP	D-form	-	-	-	0.25
	方叶无褶苔 Leiocolea bantriensis	LB	D-form	-	-	-	0.25
挺叶苔科 Anastrophyllaceae	小挺叶苔 Anastrophyllum minutum	AM	R-mat	-	-	-	0.37
裂叶苔科 Lophoziaceae	三瓣苔 Tritomaria exsecta	TE	T-form	-	0.85	-	0.62
折叶苔科 Scapaniaceae	刺边合叶苔 Scapania ciliata	SC	T-form	-	-	1.27	-
睫毛苔科 Blepharostomataceae	睫毛苔 Blepharostoma trichophyllum	BT	R-mat	-	0.50	0.08	1.37
指叶苔科 Lepidoziaceae	指叶苔 Lepidozia reptans	LR	R-mat	-	-	-	0.62
光萼苔科 Porellaceae	光萼苔 Porella pinnata	PP	R-mat	0.19	-	-	-
耳叶苔科 Frullaniaceae	细茎耳叶苔 Frullania bolanderi	FB	T-form	0.19	-	-	-
耳叶苔科 Frullaniaceae	达乌里耳叶苔 Frullania davurica	FD	T-form	1.27	-	0.64	-
	列胞耳叶苔 Frullania moniliata	FM	T-form	0.49	-	-	-
细鳞苔科 Lejeuneaceae	角齿细鳞苔 Lejeunea otiana	LO	T-form	-	-	-	0.50
细鳞苔科 Lejeuneaceae	南亚瓦鳞苔 Trocholejeunea sandvicensis	TS	T-form	0.49	-	-	-

科 Family	物种 Species	代码 Code	生活型 Life form	相对盖度 Relative coverage (%)
科 Family	物种 Species	代码 Code	生活型 Life form	Log 1	Log 2	Log 3	Log 4
曲背藓科 Oncophoraceae	曲背藓 Oncophorus wahlenbergii	OW	T-turf	9.74	-	-	-
曲背藓科 Oncophoraceae	南亚合睫藓 Symblepharis reinwardtii	SR	T-turf	-	-	-	1.99
	合睫藓 Symblepharis vaginata	SV	T-turf	-	44.18	-	2.99
曲尾藓科 Dicranaceae	波叶曲尾藓 Dicranum polysetum	DP	T-turf	-	-	3.97	-
白发藓科 Leucobryaceae	丛叶青毛藓 Dicranodontium caespitosum	DC	T-turf	-	8.52	4.77	6.97
	青毛藓 Dicranodontium denudatum	DiDe	T-turf	-	6.39	-	36.82
	长叶青毛藓 Dicranodontium didymodon	DiDi	T-turf	-	3.69	0.64	5.47
丛藓科 Pottiaceae	高山红叶藓 Bryoerythrophyllum alpigenum	BA	S-turf	-	-	0.32	-
	齿叶薄齿藓 Leptodontium handelii	LH	S-turf	0.49	-	-	-
	狭叶拟合睫藓 Pseudosymblepharis angustata	PA	S-turf	-	0.14	4.45	4.23
真藓科 Bryaceae	拟大叶真藓 Bryum salakense	BryS	T-turf	-	-	0.32	-
提灯藓科 Mniaceae	侧枝匐灯藓 Plagiomnium maximoviczii	PM	L-cushion	-	-	8.10	-
	全缘匐灯藓 Plagiomnium integrum	PI	L-cushion	-	-	-	1.00
	钝叶匐灯藓 Plagiomnium rostratum	PR	L-cushion	-	-	7.15	-
	具缘提灯藓 Mnium marginatum	MM	L-cushion	-	-	2.54	-
木灵藓科 Orthotrichaceae	毛帽木灵藓 Orthotrichum dasymitrium	OD	D-form	13.92	-	-	-
白齿藓科 Leucodontaceae	中华白齿藓 Leucodon sinensis	LS	Weft	11.68	-	-	-
蔓藓科 Meteoriaceae	新丝藓 Neodicladiella pendula	NP	R-mat	11.68	-	-	-
羽藓科 Thuidiaceae	毛羽藓 Bryonoguchia molkenboeri	BM	R-mat	-	-	-	0.50
羽藓科 Thuidiaceae	大羽藓 Thuidium cymbifolium	TC	R-mat	-	12.07	34.95	9.70
	毛尖羽藓 Thuidium plumulosum	TP	R-mat	-	-	-	0.50
	狭叶小羽藓 Haplocladium angustifolium	HA	R-mat	-	0.28	-	-
	细叶小羽藓 Haplocladium microphyllum	HM	R-mat	-	-	8.26	5.22
青藓科 Brachytheciaceae	多褶青藓 Brachythecium buchananii	BB	L-cushion	-	0.21	6.04	0.50
青藓科 Brachytheciaceae	平枝青藓 Brachythecium helminthocladum	BH	L-cushion	-	-	2.38	-
	燕尾青藓 Brachythecium novae-angliae	BraN	L-cushion	-	-	4.29	-
	褶叶青藓 Brachythecium salebrosum	BraS	L-cushion	-	-	-	1.24
	拟异叶藓 Pseudokindbergia dumosa	PD	L-cushion	-	-	0.64	-
灰藓科 Hypnaceae	圆枝粗枝藓 Gollania tereticaulis	GT	R-mat	-	2.13	-	-
灰藓科 Hypnaceae	灰藓 Hypnum cupressiforme	HC	R-mat	-	-	2.22	0.50
	密枝灰藓 Hypnum densirameum	HD	R-mat	-	2.13	2.86	7.46
	毛梳藓 Ptilium crista-castrensis	PC	R-mat	14.61	-	-	-
	拟硬叶藓 Stereodontopsis pseudorevoluta	SP	R-mat	0.19	-	-	-
	丝金灰藓 Pylaisia levieri	PL	R-mat	34.08	7.39	-	-
毛锦藓科 Pylaisiadelphaceae	曲叶小锦藓 Brotherella curvirostris	BC	R-mat	-	2.70	1.59	2.24
毛锦藓科 Pylaisiadelphaceae	弯叶小锦藓 Brotherella falcata	BF	R-mat	0.97	-	-	-
	垂蒴小锦藓 Brotherella nictans	BroN	R-mat	-	-	2.38	7.46
	暗绿毛锦藓 Pylaisiadelpha tristoviridis	PT	R-mat	-	4.12	-	-
叶苔科 Jungermanniaceae	深绿叶苔 Jungermannia atrovirens	JA	D-form	-	4.69	0.16	1.24
叶苔科 Jungermanniaceae	小萼叶苔 Jungermannia parviperiantha	JP	D-form	-	-	-	0.25
	方叶无褶苔 Leiocolea bantriensis	LB	D-form	-	-	-	0.25
挺叶苔科 Anastrophyllaceae	小挺叶苔 Anastrophyllum minutum	AM	R-mat	-	-	-	0.37
裂叶苔科 Lophoziaceae	三瓣苔 Tritomaria exsecta	TE	T-form	-	0.85	-	0.62
折叶苔科 Scapaniaceae	刺边合叶苔 Scapania ciliata	SC	T-form	-	-	1.27	-
睫毛苔科 Blepharostomataceae	睫毛苔 Blepharostoma trichophyllum	BT	R-mat	-	0.50	0.08	1.37
指叶苔科 Lepidoziaceae	指叶苔 Lepidozia reptans	LR	R-mat	-	-	-	0.62
光萼苔科 Porellaceae	光萼苔 Porella pinnata	PP	R-mat	0.19	-	-	-
耳叶苔科 Frullaniaceae	细茎耳叶苔 Frullania bolanderi	FB	T-form	0.19	-	-	-
耳叶苔科 Frullaniaceae	达乌里耳叶苔 Frullania davurica	FD	T-form	1.27	-	0.64	-
	列胞耳叶苔 Frullania moniliata	FM	T-form	0.49	-	-	-
细鳞苔科 Lejeuneaceae	角齿细鳞苔 Lejeunea otiana	LO	T-form	-	-	-	0.50
细鳞苔科 Lejeuneaceae	南亚瓦鳞苔 Trocholejeunea sandvicensis	TS	T-form	0.49	-	-	-

组合名称 Society name	物种组成 Species composition	重要值 Important value	分布样方 Distribution sample	组合名称 Society name	物种组成 Species composition	重要值 Important value	分布样方 Distribution sample
合睫藓组合 Symblepharis vaginata society	SV	0.425	Log 2-4, Log 2-5, Log 2-6, Log 2-7, Log 2-8, Log 2-9, Log 2-10	丛叶青毛藓组合 Dicranodontium vaginata society	DC	0.250	Log 3-10, Log 4-7
	DC	0.135			PA	0.167
	JA	0.100			BC	0.114
	BC	0.076			JA	0.092
	PL	0.065			PM	0.084
	DiDi	0.043			TC	0.076
	PT	0.043			BroN	0.069
	DiDe	0.034			SV	0.053
	HD	0.034			TP	0.053
	TE	0.025			BT	0.042
	BT	0.020		侧枝匐灯藓组合 Plagiomnium maximoviczii society	PM	0.346	Log 3-5, Log 3-7
大羽藓组合 Thuidium cymbifolium society	TC	0.423	Log 2-2, Log 3-2, Log 3-3, Log 3-8, Log 3-9, Log 4-10		BB	0.207
	DC	0.069			TC	0.164
	BroN	0.069			PA	0.126
	HD	0.052			HC	0.086
	PM	0.048			BraN	0.071
	DP	0.042		丝金灰藓组合 Pylaisia levieri society	PL	0.427	Log 1-1, Log 1-7, Log 1-9
	PA	0.042			PC	0.172
	JA	0.037			OW	0.132
	BT	0.036			OD	0.058
	BH	0.032			LH	0.054
	BB	0.027			FD	0.054
	SC	0.025			TS	0.052
	PT	0.021			FB	0.051
	LR	0.018		密枝灰藓组合 Hypnum densirameum society	HD	0.389	Log 4-4, Log 4-8
	SV	0.018			DiDe	0.130
	LB	0.018			BC	0.130
	DiDi	0.018			TC	0.093
青毛藓组合 Dicranodontium denudatum society	DiDe	0.391	Log 2-3, Log 4-2, Log 4-5, Log 4-6, Log 4-9		DiDi	0.074
	SR	0.064			BT	0.065
	PL	0.062			TE	0.060
	BT	0.056			LR	0.060
	JA	0.051		钝叶匐灯藓组合 Plagiomnium rostratum society	PR	0.575	Log 3-6
	GT	0.044			HM	0.145
	BroN	0.044			BB	0.145
	SV	0.035			TC	0.135
	PA	0.035		燕尾青藓组合 Brachythecium novae-angliae society	BraN	0.378	Log 3-4
	BraS	0.026			HC	0.211
	HC	0.020			TC	0.156
	BM	0.020			PA	0.133
	TE	0.020			BryS	0.122
	BB	0.020		毛帽木灵藓组合 Orthotrichum dasymitrium society	OD	0.509	Log 1-4, Log 1-5, Log 1-10
	AM	0.019			FD	0.139
	HM	0.018			TS	0.086
	LB	0.018			PL	0.068
	BC	0.018			BF	0.068
	LR	0.018			PP	0.044
	JP	0.018			SP	0.044
细叶小羽藓组合 Haplocladium microphylllum society	HM	0.282	Log 3-1, Log 4-1		FB	0.041
	DiDi	0.136		新丝藓组合 Neodicladiella pendula society	NP	0.730	Log 1-8
	PA	0.091			FM	0.270
	MM	0.081
	SV	0.061
	BB	0.049		中华白齿藓组合 Leucodon sinensis society	LS	1.000	Log 1-2
	PI	0.042
	PD	0.042
	FD	0.042		狭叶小羽藓组合 Haplocladium angustifolium society	HA	0.389	Log 2-1
	BA	0.036			BB	0.333
	LO	0.036			PA	0.278
	TC	0.036
	BT	0.036
	LR	0.031

色季拉山不同腐解等级华山松倒木上的苔藓植物组合

Bryophyte societies on the fallen logs of Pinus armandii with different decay classes in Sygera Mountains

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物种重要值之和 Sum of important values of species	大垫状 Large cushion	矮藓丛状 Short turf	直立高藓丛状 Tall turf	毯状 Rough mat	交织状 Weft	丝状 Thread-like form	紧密簇状 Dense form	组合空间结构 Society spatial structure
合睫藓组合 Symblepharis vaginata society	-	-	0.637	0.238	-	0.025	0.100	较简单 Relatively simple
大羽藓组合 Thuidium cymbifolium society	0.108	0.042	0.148	0.621	-	0.025	0.055	复杂 Complex
青毛藓组合 Dicranodontium denudatum society	0.046	0.035	0.491	0.321	-	0.020	0.088	复杂 Complex
细叶小羽藓组合 Haplocladium microphyllum society	0.213	0.091	0.233	0.385	-	0.078	-	较复杂 Relatively complex
丛叶青毛藓组合 Dicranodontium caespitosum society	0.084	0.167	0.303	0.354	-	-	0.092	较复杂 Relatively complex
侧枝匐灯藓组合 Plagiomnium maximoviczii society	0.624	0.126	-	0.250	-	-	-	较简单 Simple
丝金灰藓组合 Pylaisia levieri society	-	0.054	0.132	0.600	-	0.157	0.058	较复杂 Relatively complex
密枝灰藓组合 Hypnum densirameum society	-	-	0.204	0.736	-	0.060	-	较简单 Relatively simple
钝叶匐灯藓组合 Plagiomnium rostratum society	0.720	-	-	0.28	-	-	-	简单 Simple
燕尾青藓组合 Brachythecium novae-angliae society	-	-	-	0.225	-	0.266	0.509	较简单 Relatively simple
毛帽木灵藓组合 Orthotrichum dasymitrium society	0.378	0.133	0.122	0.367	-	-	-	较简单 Relatively simple
新丝藓组合 Neodicladiella pendula society	-	-	-	0.730	-	0.270	-	简单 Simple
中华白齿藓组合 Leucodon sinensis society	-	-	-	-	1.00	-	-	简单 Simple
狭叶小羽藓组合 Haplocladium angustifolium society	0.333	0.278	-	0.389	-	-	-	较简单 Relatively simple

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