植物生态学报 ›› 2008, Vol. 32 ›› Issue (4): 825-837.DOI: 10.3773/j.issn.1005-264x.2008.04.011
收稿日期:
2007-09-13
接受日期:
2008-01-11
出版日期:
2008-09-13
发布日期:
2008-07-30
通讯作者:
江明喜
作者简介:
*E-mail:mxjiang@wbgcas.cn基金资助:
WEI Xin-Zeng1,2, HUANG Han-Dong1, JIANG Ming-Xi1,*(), YANG Jing-Yuan3
Received:
2007-09-13
Accepted:
2008-01-11
Online:
2008-09-13
Published:
2008-07-30
Contact:
JIANG Ming-Xi
摘要:
东亚特有珍稀濒危植物领春木(Euptelea pleiospermum)在神农架地区集中分布于河岸带中。为初步探讨领春木在河岸带集中分布的成因和维持机制, 在神农架地区4条河流——沿渡河、香溪河、南河和堵河的河岸带, 共设置了30个样方(20 m × 30 m), 从种群的大小级结构、静态生命表、存活曲线、空间分布格局及其动态5个方面分析了神农架地区河岸带中领春木种群的数量特征与空间分布格局。结果表明: 1) 幼龄期个体缺乏, 中龄期个体相对丰富, 老龄期个体数量稀少。纺锤型的大小级结构表明种群属于衰退型, 但是以萌蘖为主要更新方式使其种群在较长一段时间内得以维持; 2)现存的Ⅰ、Ⅱ级个体数少, 导致静态生命表中Ⅰ、Ⅱ级个体死亡率出现负值, 自疏现象造成第Ⅳ级个体出现死亡率高峰, 由于接近实际寿命, 在第Ⅶ级死亡率达到最高; 3) 虽然其幼苗存活率较低, 但由于其幼树的存活率较高, 其存活曲线接近DeeveyⅠ型, 表明该地区河岸带的环境条件较适宜领春木种群的生长; 4) 空间分布格局总体为聚集分布, 这与大多数珍稀植物种群一致。人为干扰和自然环境影响其分布格局, 使种群由聚集分布向随机分布发展。因此使领春木种群在河岸带中沿海拔呈现“一带多岛”现象; 5) 从幼龄期到中龄期再到老龄期, 不同发育阶段的领春木种群的分布格局由聚集分布逐渐变为随机分布。但由于分布格局受种群本身生物学特性、自然环境因素(如坡向和海拔)和人为干扰的影响, 领春木在不同河岸带中的分布格局有所差异。
魏新增, 黄汉东, 江明喜, 杨敬元. 神农架地区河岸带中领春木种群数量特征与空间分布格局. 植物生态学报, 2008, 32(4): 825-837. DOI: 10.3773/j.issn.1005-264x.2008.04.011
WEI Xin-Zeng, HUANG Han-Dong, JIANG Ming-Xi, YANG Jing-Yuan. QUANTITATIVE CHARACTERISTICS AND SPATIAL DISTRIBUTION PATTERNS OF EUPTELEA PLEIOSPERMUMPOPULATIONS IN RIPARIAN ZONES OF THE SHENNONGJIA AREA, CENTRAL CHINA. Chinese Journal of Plant Ecology, 2008, 32(4): 825-837. DOI: 10.3773/j.issn.1005-264x.2008.04.011
图1 领春木种群的大小级结构 a: 整个神农架地区河岸带中领春木种群的大小级结构 Total population ofE. pleiospermum along the riparian zones of the Shennongjia area b: 沿渡河河岸带中领春木种群的大小级结构 Population ofE. pleiospermumalong the riparian zone of the Yandu River c: 香溪河河岸带中领春木种群的大小级结构 Population ofE. pleiospermumalong the riparian zone of the Xiangxi River d: 南河河岸带中领春木种群的大小级结构 Population ofE. pleiospermumalong the riparian zone of the Nan River e: 堵河河岸带中领春木种群的大小级结构 Population ofE. pleiospermumalong the riparian zone of the Du River
Fig. 1 The size structure of Euptelea pleiospermumpopulation
大小级 Size class x | 存活数 No. of survival Nx | 存活数标准化Standard survival Lx | lgLx | 死亡数 Standard death at interval Dx | 死亡率Mortality Qx | 从第x大小级起超过x大小级的存活个体总数 T | 生命期望 Expecting life Ex |
---|---|---|---|---|---|---|---|
Ⅰ | 23 | 133.5 | 2.13 | - | - | 1 141.5 | 49.63 |
Ⅱ | 244 | 358.0 | 2.55 | - | - | 1 008.0 | 4.13 |
Ⅲ | 472 | 367.0 | 2.56 | 210 | 0.44 | 650.0 | 1.38 |
Ⅳ | 262 | 182.0 | 2.26 | 160 | 0.61 | 283.0 | 1.08 |
Ⅴ | 102 | 73.0 | 1.86 | 58 | 0.57 | 101.0 | 0.99 |
Ⅵ | 44 | 25.0 | 1.40 | 38 | 0.86 | 28.0 | 0.64 |
Ⅶ | 6 | 3.0 | 0.48 | 6 | 1.00 | 3.0 | 0.50 |
表1 整个神农架地区河岸带中领春木种群的静态生命表
Table 1 Static life table of Euptelea pleiospermum population along the riparian zones of the Shennongjia area
大小级 Size class x | 存活数 No. of survival Nx | 存活数标准化Standard survival Lx | lgLx | 死亡数 Standard death at interval Dx | 死亡率Mortality Qx | 从第x大小级起超过x大小级的存活个体总数 T | 生命期望 Expecting life Ex |
---|---|---|---|---|---|---|---|
Ⅰ | 23 | 133.5 | 2.13 | - | - | 1 141.5 | 49.63 |
Ⅱ | 244 | 358.0 | 2.55 | - | - | 1 008.0 | 4.13 |
Ⅲ | 472 | 367.0 | 2.56 | 210 | 0.44 | 650.0 | 1.38 |
Ⅳ | 262 | 182.0 | 2.26 | 160 | 0.61 | 283.0 | 1.08 |
Ⅴ | 102 | 73.0 | 1.86 | 58 | 0.57 | 101.0 | 0.99 |
Ⅵ | 44 | 25.0 | 1.40 | 38 | 0.86 | 28.0 | 0.64 |
Ⅶ | 6 | 3.0 | 0.48 | 6 | 1.00 | 3.0 | 0.50 |
大小级 Size class x | 存活数No. of survival Nx | 存活数标准化Standard survival Lx | lgLx | 死亡数 Standard death at interval Dx | 死亡率Mortality Qx | 从第x大小级起超过x大小级的存活个体总数 T | 生命期望 Expecting life Ex |
---|---|---|---|---|---|---|---|
Ⅰ | 20 | 58.0 | 1.76 | - | - | 343.0 | 17.15 |
Ⅱ | 96 | 113.5 | 2.05 | - | - | 285.0 | 2.97 |
Ⅲ | 131 | 96.5 | 1.98 | 69 | 0.53 | 171.5.0 | 1.31 |
Ⅳ | 62 | 42.0 | 1.62 | 40 | 0.65 | 75.0 | 1.21 |
Ⅴ | 22 | 21.0 | 1.32 | 2 | 0.09 | 33.0 | 1.5.0 |
Ⅵ | 20 | 11.0 | 1.04 | 18 | 0.90 | 12.0 | 0.60 |
Ⅶ | 2 | 1.0 | 0 | 2 | 1.00 | 1.0 | 0.50 |
表2 沿渡河河岸带中领春木种群的静态生命表
Table 2 Static life table of Euptelea pleiospermum population along the riparian zone of the Yandu River
大小级 Size class x | 存活数No. of survival Nx | 存活数标准化Standard survival Lx | lgLx | 死亡数 Standard death at interval Dx | 死亡率Mortality Qx | 从第x大小级起超过x大小级的存活个体总数 T | 生命期望 Expecting life Ex |
---|---|---|---|---|---|---|---|
Ⅰ | 20 | 58.0 | 1.76 | - | - | 343.0 | 17.15 |
Ⅱ | 96 | 113.5 | 2.05 | - | - | 285.0 | 2.97 |
Ⅲ | 131 | 96.5 | 1.98 | 69 | 0.53 | 171.5.0 | 1.31 |
Ⅳ | 62 | 42.0 | 1.62 | 40 | 0.65 | 75.0 | 1.21 |
Ⅴ | 22 | 21.0 | 1.32 | 2 | 0.09 | 33.0 | 1.5.0 |
Ⅵ | 20 | 11.0 | 1.04 | 18 | 0.90 | 12.0 | 0.60 |
Ⅶ | 2 | 1.0 | 0 | 2 | 1.00 | 1.0 | 0.50 |
大小级 Size class x | 存活数No. of survival Nx | 存活数标准化Standard survival Lx | lgLx | 死亡数 Standard death at interval Dx | 死亡率Mortality Qx | 从第x大小级起超过x大小级的存活个体总数 T | 生命期望 Expecting life Ex |
---|---|---|---|---|---|---|---|
Ⅰ | 3 | 34.5 | 1.54 | - | - | 395.5 | 131.83 |
Ⅱ | 66 | 132.5 | 2.12 | - | - | 361.0 | 5.47 |
Ⅲ | 199 | 153.0 | 2.18 | 92 | 0.46 | 228.5 | 1.15 |
Ⅳ | 107 | 62.5 | 1.80 | 89 | 0.83 | 75.5 | 0.71 |
Ⅴ | 18 | 11.0 | 1.04 | 14 | 0.78 | 13.0 | 0.72 |
Ⅵ | 4 | 2.0 | 0.30 | 4 | 1.00 | 2.0 | 0.50 |
Ⅶ | 0 | 0 | - | 0 | - | 0 | - |
表3 香溪河河岸带中领春木种群的静态生命表
Table 3 Static life table of Euptelea pleiospermum population along the riparian zone of the Xiangxi River
大小级 Size class x | 存活数No. of survival Nx | 存活数标准化Standard survival Lx | lgLx | 死亡数 Standard death at interval Dx | 死亡率Mortality Qx | 从第x大小级起超过x大小级的存活个体总数 T | 生命期望 Expecting life Ex |
---|---|---|---|---|---|---|---|
Ⅰ | 3 | 34.5 | 1.54 | - | - | 395.5 | 131.83 |
Ⅱ | 66 | 132.5 | 2.12 | - | - | 361.0 | 5.47 |
Ⅲ | 199 | 153.0 | 2.18 | 92 | 0.46 | 228.5 | 1.15 |
Ⅳ | 107 | 62.5 | 1.80 | 89 | 0.83 | 75.5 | 0.71 |
Ⅴ | 18 | 11.0 | 1.04 | 14 | 0.78 | 13.0 | 0.72 |
Ⅵ | 4 | 2.0 | 0.30 | 4 | 1.00 | 2.0 | 0.50 |
Ⅶ | 0 | 0 | - | 0 | - | 0 | - |
大小级 Size class x | 存活数 No. of survival Nx | 存活数标准化Standard survival Lx | lgLx | 死亡数 Standard death at interval Dx | 死亡率Mortality Qx | 从第x大小级起超过x大小级的存活个体总数 T | 生命期望 Expecting life Ex |
---|---|---|---|---|---|---|---|
Ⅰ | 2 | 30.0 | 1.48 | - | - | 234.0 | 117.00 |
Ⅱ | 58 | 78.5 | 1.89 | - | - | 204.0 | 3.52 |
Ⅲ | 99 | 79.0 | 1.90 | 40 | 0.40 | 125.5 | 1.27 |
Ⅳ | 59 | 37.5 | 1.57 | 43 | 0.73 | 46.5 | 0.79 |
Ⅴ | 16 | 8.5 | 0.93 | 15 | 0.94 | 9.0 | 0.56 |
Ⅵ | 1 | 0.5 | -0.30 | 1 | 1.00 | 0.5 | 0.50 |
Ⅶ | 0 | 0 | - | 0 | - | 0 | - |
表4 南河河岸带中领春木种群的静态生命表
Table 4 Static life table of Euptelea pleiospermum population along the riparian zone of the Nan River
大小级 Size class x | 存活数 No. of survival Nx | 存活数标准化Standard survival Lx | lgLx | 死亡数 Standard death at interval Dx | 死亡率Mortality Qx | 从第x大小级起超过x大小级的存活个体总数 T | 生命期望 Expecting life Ex |
---|---|---|---|---|---|---|---|
Ⅰ | 2 | 30.0 | 1.48 | - | - | 234.0 | 117.00 |
Ⅱ | 58 | 78.5 | 1.89 | - | - | 204.0 | 3.52 |
Ⅲ | 99 | 79.0 | 1.90 | 40 | 0.40 | 125.5 | 1.27 |
Ⅳ | 59 | 37.5 | 1.57 | 43 | 0.73 | 46.5 | 0.79 |
Ⅴ | 16 | 8.5 | 0.93 | 15 | 0.94 | 9.0 | 0.56 |
Ⅵ | 1 | 0.5 | -0.30 | 1 | 1.00 | 0.5 | 0.50 |
Ⅶ | 0 | 0 | - | 0 | - | 0 | - |
大小级 Size class x | 存活数 No. of survival Nx | 存活数标准化Standard survival Lx | lgLx | 死亡数 Standard death at interval Dx | 死亡率Mortality Qx | 从第x大小级起超过x大小级的存活个体总数 T | 生命期望 Expecting life Ex |
---|---|---|---|---|---|---|---|
Ⅰ | 0 | 12.0 | 1.08 | - | - | 170.0 | - |
Ⅱ | 24 | 33.5 | 1.53 | - | - | 158.0 | 6.58 |
Ⅲ | 43 | 38.5 | 1.59 | 9 | 0.21 | 124.5 | 2.90 |
Ⅳ | 34 | 40.0 | 1.60 | - | - | 86.0 | 2.53 |
Ⅴ | 46 | 32.5 | 1.51 | 27 | 0.59 | 46.0 | 1.00 |
Ⅵ | 19 | 11.5 | 1.06 | 15 | 0.79 | 13.5 | 0.71 |
Ⅶ | 4 | 2.0 | 0.30 | 4 | 1.00 | 2.0 | 0.50 |
表5 堵河河岸带中领春木种群的静态生命表
Table 5 Static life table of Euptelea pleiospermum population along the riparian zone of the Du River
大小级 Size class x | 存活数 No. of survival Nx | 存活数标准化Standard survival Lx | lgLx | 死亡数 Standard death at interval Dx | 死亡率Mortality Qx | 从第x大小级起超过x大小级的存活个体总数 T | 生命期望 Expecting life Ex |
---|---|---|---|---|---|---|---|
Ⅰ | 0 | 12.0 | 1.08 | - | - | 170.0 | - |
Ⅱ | 24 | 33.5 | 1.53 | - | - | 158.0 | 6.58 |
Ⅲ | 43 | 38.5 | 1.59 | 9 | 0.21 | 124.5 | 2.90 |
Ⅳ | 34 | 40.0 | 1.60 | - | - | 86.0 | 2.53 |
Ⅴ | 46 | 32.5 | 1.51 | 27 | 0.59 | 46.0 | 1.00 |
Ⅵ | 19 | 11.5 | 1.06 | 15 | 0.79 | 13.5 | 0.71 |
Ⅶ | 4 | 2.0 | 0.30 | 4 | 1.00 | 2.0 | 0.50 |
河流 River | 海拔 Altitude (m) | 方差均值比 v/ | 平均拥挤度 Index of mean crowding | 聚块性指数 Patchiness index | Morisita指数 Morisita index | 格局 Pattern | ||
---|---|---|---|---|---|---|---|---|
Value | t | Value | F | |||||
沿渡河 Yandu River | 900 | 4.29 | 11.16*** | 5.50 | 2.49 | 2.46 | 4.29** | C |
1 000 | 3.03 | 6.88*** | 2.94 | 3.21 | 3.22 | 3.03** | C | |
1 200 | 1.86 | 2.91* | 1.82 | 1.90 | 1.90 | 1.86* | C | |
1 300 | 1.49 | 1.67 | 1.62 | 1.44 | 1.44 | 1.49 | R | |
1 400 | 1.50 | 1.68 | 1.45 | 1.52 | 1.52 | 1.50 | R | |
1 500 | 4.10 | 10.53*** | 5.23 | 2.46 | 2.43 | 4.10** | C | |
1 600 | 2.42 | 4.80** | 3.13 | 1.83 | 1.81 | 2.42** | C | |
1 700 | 1.04 | 0.14 | 0.75 | 1.06 | 1.06 | 1.04 | R | |
1 800 | 1.22 | 0.74 | 0.55 | 1.65 | 1.71 | 1.22 | R | |
1 900 | 1.38 | 1.28 | 1.67 | 1.29 | 1.29 | 1.38 | R | |
2 000 | 1.84 | 2.85* | 3.13 | 1.37 | 1.36 | 1.84* | C | |
香溪河 Xiangxi River | 1 100 | 2.32 | 4.47** | 2.19 | 2.50 | 2.51 | 2.32** | C |
1 300 | 2.86 | 6.30*** | 2.81 | 2.94 | 2.94 | 2.86** | C | |
1 400 | 2.61 | 5.48*** | 2.49 | 2.85 | 2.86 | 2.61** | C | |
1 500 | 1.43 | 1.44 | 1.26 | 1.51 | 1.52 | 1.43 | R | |
1 600 | 2.30 | 4.41** | 2.51 | 2.08 | 2.07 | 2.30** | C | |
1 700 | 2.59 | 5.39*** | 2.80 | 2.32 | 2.31 | 2.59** | C | |
1 800 | 1.13 | 0.42 | 1.17 | 1.12 | 1.12 | 1.13 | R | |
1 900 | 1.30 | 1.03 | 1.05 | 1.41 | 1.41 | 1.30 | R | |
南河 Nan Rvier | 1 300 | 3.67 | 9.11*** | 3.77 | 3.48 | 3.47 | 3.69** | C |
1 400 | 2.08 | 3.66** | 2.00 | 2.18 | 2.18 | 2.08** | C | |
1 500 | 2.61 | 5.46*** | 3.11 | 2.07 | 2.13 | 2.72** | C | |
1 600 | 2.34 | 4.54** | 1.96 | 3.14 | 3.20 | 2.34** | C | |
1 700 | 2.95 | 6.60*** | 2.91 | 3.03 | 3.04 | 2.95** | C | |
1 800 | 4.22 | 10.91*** | 4.18 | 4.36 | 3.89 | 3.76** | C | |
1 900 | 6.57 | 18.88*** | 8.11 | 3.19 | 3.13 | 6.57*** | C | |
堵河 Du Rvier | 1 700 | 3.09 | 7.08*** | 2.75 | 4.13 | 4.20 | 3.09** | C |
1 800 | 1.79 | 2.69* | 1.83 | 1.76 | 1.76 | 1.79* | C | |
1 900 | 1.32 | 1.09 | 1.16 | 1.39 | 1.39 | 1.32 | R | |
2 000 | 1.42 | 1.41 | 1.33 | 1.45 | 1.45 | 1.42 | R |
表6 神农架地区河岸带中领春木种群沿海拔的空间分布格局
Table 6 Spatial distribution patterns of Euptelea pleiospermumpopulation along the riparian zones at different altitudes
河流 River | 海拔 Altitude (m) | 方差均值比 v/ | 平均拥挤度 Index of mean crowding | 聚块性指数 Patchiness index | Morisita指数 Morisita index | 格局 Pattern | ||
---|---|---|---|---|---|---|---|---|
Value | t | Value | F | |||||
沿渡河 Yandu River | 900 | 4.29 | 11.16*** | 5.50 | 2.49 | 2.46 | 4.29** | C |
1 000 | 3.03 | 6.88*** | 2.94 | 3.21 | 3.22 | 3.03** | C | |
1 200 | 1.86 | 2.91* | 1.82 | 1.90 | 1.90 | 1.86* | C | |
1 300 | 1.49 | 1.67 | 1.62 | 1.44 | 1.44 | 1.49 | R | |
1 400 | 1.50 | 1.68 | 1.45 | 1.52 | 1.52 | 1.50 | R | |
1 500 | 4.10 | 10.53*** | 5.23 | 2.46 | 2.43 | 4.10** | C | |
1 600 | 2.42 | 4.80** | 3.13 | 1.83 | 1.81 | 2.42** | C | |
1 700 | 1.04 | 0.14 | 0.75 | 1.06 | 1.06 | 1.04 | R | |
1 800 | 1.22 | 0.74 | 0.55 | 1.65 | 1.71 | 1.22 | R | |
1 900 | 1.38 | 1.28 | 1.67 | 1.29 | 1.29 | 1.38 | R | |
2 000 | 1.84 | 2.85* | 3.13 | 1.37 | 1.36 | 1.84* | C | |
香溪河 Xiangxi River | 1 100 | 2.32 | 4.47** | 2.19 | 2.50 | 2.51 | 2.32** | C |
1 300 | 2.86 | 6.30*** | 2.81 | 2.94 | 2.94 | 2.86** | C | |
1 400 | 2.61 | 5.48*** | 2.49 | 2.85 | 2.86 | 2.61** | C | |
1 500 | 1.43 | 1.44 | 1.26 | 1.51 | 1.52 | 1.43 | R | |
1 600 | 2.30 | 4.41** | 2.51 | 2.08 | 2.07 | 2.30** | C | |
1 700 | 2.59 | 5.39*** | 2.80 | 2.32 | 2.31 | 2.59** | C | |
1 800 | 1.13 | 0.42 | 1.17 | 1.12 | 1.12 | 1.13 | R | |
1 900 | 1.30 | 1.03 | 1.05 | 1.41 | 1.41 | 1.30 | R | |
南河 Nan Rvier | 1 300 | 3.67 | 9.11*** | 3.77 | 3.48 | 3.47 | 3.69** | C |
1 400 | 2.08 | 3.66** | 2.00 | 2.18 | 2.18 | 2.08** | C | |
1 500 | 2.61 | 5.46*** | 3.11 | 2.07 | 2.13 | 2.72** | C | |
1 600 | 2.34 | 4.54** | 1.96 | 3.14 | 3.20 | 2.34** | C | |
1 700 | 2.95 | 6.60*** | 2.91 | 3.03 | 3.04 | 2.95** | C | |
1 800 | 4.22 | 10.91*** | 4.18 | 4.36 | 3.89 | 3.76** | C | |
1 900 | 6.57 | 18.88*** | 8.11 | 3.19 | 3.13 | 6.57*** | C | |
堵河 Du Rvier | 1 700 | 3.09 | 7.08*** | 2.75 | 4.13 | 4.20 | 3.09** | C |
1 800 | 1.79 | 2.69* | 1.83 | 1.76 | 1.76 | 1.79* | C | |
1 900 | 1.32 | 1.09 | 1.16 | 1.39 | 1.39 | 1.32 | R | |
2 000 | 1.42 | 1.41 | 1.33 | 1.45 | 1.45 | 1.42 | R |
河流 River | 海拔 Altitude (m) | 阶段 Period | 方差均值比 v/ | 平均拥挤度 Index of mean crowding | 聚块性指数 Patchiness index | Morisita指数 Morisita index | 格局 Pattern | ||
---|---|---|---|---|---|---|---|---|---|
Value | t | Value | F | ||||||
沿渡河 Yandu River | 900 | Y | 4.45 | 11.70*** | 5.49 | 2.69 | 2.65 | 4.45** | C |
M | 0.91 | -0.29 | 0.04 | 0.30 | 0.00 | 0.91 | R | ||
O | 1.24 | 0.83 | 0.45 | 2.17 | 2.40 | 1.24 | R | ||
1 400 | Y | 2.51 | 5.11*** | 1.88 | 5.02 | 5.33 | 2.51** | C | |
M | 0.78 | -0.74 | 0.03 | 0.13 | 0.00 | 0.78 | R | ||
O | 0.96 | -0.15 | 0.29 | 0.87 | 0.86 | 0.96 | R | ||
1 600 | Y | 1.89 | 3.03** | 1.35 | 2.95 | 3.05 | 1.89* | C | |
M | 1.63 | 2.12* | 1.67 | 1.60 | 1.60 | 1.63* | C | ||
O | 1.48 | 1.62 | 0.73 | 2.91 | 3.20 | 1.48 | R | ||
1 900 | Y | 1.24 | 0.83 | 0.45 | 2.17 | 2.40 | 1.24 | R | |
M | 1.04 | 0.14 | 0.75 | 1.06 | 1.06 | 1.04 | R | ||
O | 1.13 | 0.46 | 0.59 | 1.29 | 1.31 | 1.13 | R | ||
香溪河 Xiangxi River | 1 100 | Y | 2.04 | 3.54** | 1.42 | 3.78 | 4.00 | 2.04** | C |
M | 2.09 | 3.69** | 1.59 | 3.17 | 3.27 | 2.09** | C | ||
O | 1.00 | 0.00 | 0.04 | 1.00 | - | - | R | ||
1 500 | Y | - | - | - | - | - | - | N | |
M | 1.32 | 1.07 | 1.11 | 1.40 | 1.40 | 1.32 | R | ||
O | 1.00 | 0.00 | 0.04 | 1.00 | - | - | R | ||
1 700 | Y | - | - | - | - | - | - | N | |
M | 2.65 | 5.61*** | 3.32 | 1.99 | 2.31 | 2.59** | C | ||
O | - | - | - | - | - | - | N | ||
1 900 | Y | 1.00 | 0.00 | 0.04 | 1.00 | - | - | R | |
M | 1.16 | 0.56 | 0.87 | 1.23 | 1.24 | 1.16 | R | ||
O | 1.61 | 2.06* | 0.73 | 5.87 | 8.00 | 1.61* | C | ||
南河 Nan River | 1 300 | Y | 4.00 | 10.17*** | 4.00 | 4.00 | 4.00 | 4.00** | C |
M | 1.24 | 0.83 | 0.45 | 2.17 | 2.40 | 1.24 | R | ||
O | - | - | - | - | - | - | N | ||
1 600 | Y | 0.91 | -0.29 | 0.04 | 0.30 | 0.00 | 0.91 | R | |
M | 2.61 | 5.46*** | 2.11 | 4.22 | 4.36 | 2.61** | C | ||
O | - | - | - | - | - | - | N | ||
1 900 | Y | 0.96 | -0.15 | 0.29 | 0.87 | 0.86 | 0.96 | R | |
M | 6.26 | 17.84*** | 7.47 | 3.38 | 3.33 | 6.26*** | C | ||
O | - | - | - | - | - | - | N | ||
堵河 Du River | 1 700 | Y | 1.00 | 0.00 | 0.04 | 1.00 | - | - | R |
M | 2.52 | 5.16*** | 2.11 | 3.61 | 3.69 | 2.52** | C | ||
O | 1.00 | 0.00 | 0.04 | 1.00 | - | - | R | ||
1 900 | Y | 0.96 | -0.15 | 0.04 | 0.48 | 0.00 | 0.96 | R | |
M | 0.82 | -0.62 | 0.23 | 0.56 | 0.53 | 0.82 | R | ||
O | 1.12 | 0.39 | 0.49 | 1.31 | 1.33 | 1.12 | R |
表7 神农架地区河岸带中不同发育阶段的领春木种群沿海拔的空间分布格局
Table 7 Spatial distributional patterns of Euptelea pleiospermumpopulations along the riparian zones in different growth periods
河流 River | 海拔 Altitude (m) | 阶段 Period | 方差均值比 v/ | 平均拥挤度 Index of mean crowding | 聚块性指数 Patchiness index | Morisita指数 Morisita index | 格局 Pattern | ||
---|---|---|---|---|---|---|---|---|---|
Value | t | Value | F | ||||||
沿渡河 Yandu River | 900 | Y | 4.45 | 11.70*** | 5.49 | 2.69 | 2.65 | 4.45** | C |
M | 0.91 | -0.29 | 0.04 | 0.30 | 0.00 | 0.91 | R | ||
O | 1.24 | 0.83 | 0.45 | 2.17 | 2.40 | 1.24 | R | ||
1 400 | Y | 2.51 | 5.11*** | 1.88 | 5.02 | 5.33 | 2.51** | C | |
M | 0.78 | -0.74 | 0.03 | 0.13 | 0.00 | 0.78 | R | ||
O | 0.96 | -0.15 | 0.29 | 0.87 | 0.86 | 0.96 | R | ||
1 600 | Y | 1.89 | 3.03** | 1.35 | 2.95 | 3.05 | 1.89* | C | |
M | 1.63 | 2.12* | 1.67 | 1.60 | 1.60 | 1.63* | C | ||
O | 1.48 | 1.62 | 0.73 | 2.91 | 3.20 | 1.48 | R | ||
1 900 | Y | 1.24 | 0.83 | 0.45 | 2.17 | 2.40 | 1.24 | R | |
M | 1.04 | 0.14 | 0.75 | 1.06 | 1.06 | 1.04 | R | ||
O | 1.13 | 0.46 | 0.59 | 1.29 | 1.31 | 1.13 | R | ||
香溪河 Xiangxi River | 1 100 | Y | 2.04 | 3.54** | 1.42 | 3.78 | 4.00 | 2.04** | C |
M | 2.09 | 3.69** | 1.59 | 3.17 | 3.27 | 2.09** | C | ||
O | 1.00 | 0.00 | 0.04 | 1.00 | - | - | R | ||
1 500 | Y | - | - | - | - | - | - | N | |
M | 1.32 | 1.07 | 1.11 | 1.40 | 1.40 | 1.32 | R | ||
O | 1.00 | 0.00 | 0.04 | 1.00 | - | - | R | ||
1 700 | Y | - | - | - | - | - | - | N | |
M | 2.65 | 5.61*** | 3.32 | 1.99 | 2.31 | 2.59** | C | ||
O | - | - | - | - | - | - | N | ||
1 900 | Y | 1.00 | 0.00 | 0.04 | 1.00 | - | - | R | |
M | 1.16 | 0.56 | 0.87 | 1.23 | 1.24 | 1.16 | R | ||
O | 1.61 | 2.06* | 0.73 | 5.87 | 8.00 | 1.61* | C | ||
南河 Nan River | 1 300 | Y | 4.00 | 10.17*** | 4.00 | 4.00 | 4.00 | 4.00** | C |
M | 1.24 | 0.83 | 0.45 | 2.17 | 2.40 | 1.24 | R | ||
O | - | - | - | - | - | - | N | ||
1 600 | Y | 0.91 | -0.29 | 0.04 | 0.30 | 0.00 | 0.91 | R | |
M | 2.61 | 5.46*** | 2.11 | 4.22 | 4.36 | 2.61** | C | ||
O | - | - | - | - | - | - | N | ||
1 900 | Y | 0.96 | -0.15 | 0.29 | 0.87 | 0.86 | 0.96 | R | |
M | 6.26 | 17.84*** | 7.47 | 3.38 | 3.33 | 6.26*** | C | ||
O | - | - | - | - | - | - | N | ||
堵河 Du River | 1 700 | Y | 1.00 | 0.00 | 0.04 | 1.00 | - | - | R |
M | 2.52 | 5.16*** | 2.11 | 3.61 | 3.69 | 2.52** | C | ||
O | 1.00 | 0.00 | 0.04 | 1.00 | - | - | R | ||
1 900 | Y | 0.96 | -0.15 | 0.04 | 0.48 | 0.00 | 0.96 | R | |
M | 0.82 | -0.62 | 0.23 | 0.56 | 0.53 | 0.82 | R | ||
O | 1.12 | 0.39 | 0.49 | 1.31 | 1.33 | 1.12 | R |
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