基于多元回归树的常绿阔叶林群丛数量分类——以古田山24公顷森林样地为例

doi:10.3773/j.issn.1005-264x.2010.07.001

摘要/Abstract

摘要：

常绿阔叶林植被分类一直是植被生态学研究中的难题, 尤其是基本单位——群丛的分类。该文以地形因子和物种组成数据为变量的多元回归树方法, 将浙江古田山24 hm²森林监测样地的森林群落分为3个群丛。所得群丛既反映了群落在时间和空间上的相对间断分布, 也符合植被分类基本单位的特点, 为常绿阔叶林的群丛分类提供了新思路。并首次引入了指示值算法, 解决了以往指示种无法量化的难题。最终以优势层优势种为主、下木层指示种为辅的命名原则, 将3个群丛命名为: 1) 石斑木(Raphiolepis indica) + 柳叶蜡梅(Chimonanthus salicifolius)—格药柃(Eurya muricata) + 赤楠(Syzygium buxifolium)—木荷(Schima superba) + 甜槠(Castanopsis eyrei)群丛; 2) 映山红(Rhododendron simsii) + 满山红(Rhododendron mariesii)—短柄枹(Quercus serrata var. brevipetiolata) + 灰白蜡瓣花(Corylopsis glandulifera var. hypoglauca)—马尾松(Pinus massoniana) + 甜槠群丛; 3) 短尾越桔(Vaccinium carlesii) + 毛花连蕊茶(Camellia fraterna)—杨梅叶蚊母树(Distylium myricoides)+浙江新木姜子(Neolitsea aurata)—木荷+甜槠群丛。

关键词: 生境, 指示种, 指示值, 地形因子, 植被分类

Abstract:

Aims A 24 hm² permanent plot in Gutianshan National Nature Reserve provided a valuable case for association classification in evergreen broad-leaved forest. Our objectives were to divide the forest community into associations to provide a new classification of evergreen broad-leaved forest and introduce the algorithm of indicator value for species in associations. Indicator species previously could not be quantified.

Methods We used multivariate regression trees, based on topographic factors and species composition, for association classification. An indicator value was introduced to quantify the indicator species of associations. We named associations after the dominant species in the tree layer, followed by indicator species of lower tree layers.

Important findings The forest community was divided into three associations that not only reflect temporal and spatial disjunctions of the community, but also correspond with features of the basic unit of vegetation classification. The three associations are 1) Raphiolepis indica + Chimonanthus salicifolius―Eurya muricata + Syzygium buxifolium―Schima superba + Castanopsis eyrei Association; 2) Rhododendron simsii + Rhododendron mariesii―Quercus serrata var. brevipetiolata + Corylopsis glandulifera var. hypoglauca―Pinus massoniana + Castanopsis eyrei Association; and 3) Vaccinium carlesii + Camellia fraterna―Distylium myricoides + Neolitsea aurata―Schima superba + Castanopsis eyrei Association.

Key words: habitat, indicator species, indicator value, topography factor, vegetation classification

赖江山, 米湘成, 任海保, 马克平. 基于多元回归树的常绿阔叶林群丛数量分类——以古田山24公顷森林样地为例. 植物生态学报, 2010, 34(7): 761-769. DOI: 10.3773/j.issn.1005-264x.2010.07.001

LAI Jiang-Shan, MI Xiang-Cheng, REN Hai-Bao, MA Ke-Ping. Numerical classification of associations in subtropical evergreen broad-leaved forest based on multivariate regression trees―a case study of 24 hm ² Gutianshan forest plot in China. Chinese Journal of Plant Ecology, 2010, 34(7): 761-769. DOI: 10.3773/j.issn.1005-264x.2010.07.001

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链接本文: https://www.plant-ecology.com/CN/10.3773/j.issn.1005-264x.2010.07.001

https://www.plant-ecology.com/CN/Y2010/V34/I7/761

图/表 6

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物种 Species	群丛1 Asso. 1		群丛2 Asso. 2		群丛3 Asso. 3		层次 Layer
物种 Species	优势度 Dominance	排序Order	优势度 Dominance	排序order	优势度 Dominance	排序Order	层次 Layer
甜槠 Castanopsis eyrei	0.40	1	0.25	1	0.27	1	T
木荷 Schima superba	0.19	2	0.19	3	0.18	2	T
马尾松 Pinus massoniana	0.13	3	0.21	2	0.05	3	T
短柄枹 Quercus serrata var. brevipetiolata	0.04	4	0.13	4	0.01	17	U

物种 Species	群丛1 Asso. 1		群丛2 Asso. 2		群丛3 Asso. 3		层次 Layer
物种 Species	优势度 Dominance	排序Order	优势度 Dominance	排序order	优势度 Dominance	排序Order	层次 Layer
甜槠 Castanopsis eyrei	0.40	1	0.25	1	0.27	1	T
木荷 Schima superba	0.19	2	0.19	3	0.18	2	T
马尾松 Pinus massoniana	0.13	3	0.21	2	0.05	3	T
短柄枹 Quercus serrata var. brevipetiolata	0.04	4	0.13	4	0.01	17	U

群丛1 Asso. 1				群丛2 Asso. 2				群丛3 Asso. 3
物种 Species	指示值 Indicator value	层次 Layer	生活型 Life- form	物种 Species	指示值 Indicator value	层次 Layer	生活型 Life- form	物种 Species	指示值 Indicator value	层次 Layer	生活型 Life- form
赤楠 Syzygium buxifolium	48	U	E	满山红Rhododendron mariesii	71	S	D	毛花连蕊茶Camellia fraterna	62	S	E
甜槠 Castanopsis eyrei	44	T	E	灰白蜡瓣花 Corylopsis glandulifera var. hypoglauca	69	U	D	红楠 Machilus thunbergii	60	T	E
格药柃 Eurya muricata	41	U	E	映山红Rhododendron simsii	68	S	D	青冈Cyclobalanopsis glauca	57	T	E
马银花Rhododendron ovatum	40	U	E	短柄枹 Quercus serrata var. brevipetiolata	64	U	D	丝栗栲Castanopsis fargesii	53	T	E
厚皮香 Ternstroemia gymnanthera	38	U	E	马尾松 Pinus massoniana	52	T	E	香桂Cinnamomum subavenium	51	T	E
柳叶蜡梅Chimonanthus salicifolius	37	S	E	毛果南烛 Lyonia ovalifolia var. hebecarpa	47	U	D	浙江新木姜子Neolitsea aurata var. chekiangensis	50	U	E
石斑木 Raphiolepis indica	33	S	E	江南越桔Vaccinium mandarinorum	46	U	E	矩形叶鼠刺 Itea oblonga	49	U	D
豹皮樟 Litsea coreana var. sinensis	19	T	E	山合欢 Albizia kalkora	44	T	D	杨梅叶蚊母树 Distylium myricoides	48	U	E
山矾 Symplocos sumuntia	04	U	E	乌饭树 Vaccinium bracteatum	44	S	E	麂角杜鹃 Rhododendron latoucheae	46	U	E
香冬青 Ilex suaveolens	03	T	E	木荷 Schima superba	43	T	E	短尾越桔Vaccinium carlesii	44	S	E

群丛1 Asso. 1				群丛2 Asso. 2				群丛3 Asso. 3
物种 Species	指示值 Indicator value	层次 Layer	生活型 Life- form	物种 Species	指示值 Indicator value	层次 Layer	生活型 Life- form	物种 Species	指示值 Indicator value	层次 Layer	生活型 Life- form
赤楠 Syzygium buxifolium	48	U	E	满山红Rhododendron mariesii	71	S	D	毛花连蕊茶Camellia fraterna	62	S	E
甜槠 Castanopsis eyrei	44	T	E	灰白蜡瓣花 Corylopsis glandulifera var. hypoglauca	69	U	D	红楠 Machilus thunbergii	60	T	E
格药柃 Eurya muricata	41	U	E	映山红Rhododendron simsii	68	S	D	青冈Cyclobalanopsis glauca	57	T	E
马银花Rhododendron ovatum	40	U	E	短柄枹 Quercus serrata var. brevipetiolata	64	U	D	丝栗栲Castanopsis fargesii	53	T	E
厚皮香 Ternstroemia gymnanthera	38	U	E	马尾松 Pinus massoniana	52	T	E	香桂Cinnamomum subavenium	51	T	E
柳叶蜡梅Chimonanthus salicifolius	37	S	E	毛果南烛 Lyonia ovalifolia var. hebecarpa	47	U	D	浙江新木姜子Neolitsea aurata var. chekiangensis	50	U	E
石斑木 Raphiolepis indica	33	S	E	江南越桔Vaccinium mandarinorum	46	U	E	矩形叶鼠刺 Itea oblonga	49	U	D
豹皮樟 Litsea coreana var. sinensis	19	T	E	山合欢 Albizia kalkora	44	T	D	杨梅叶蚊母树 Distylium myricoides	48	U	E
山矾 Symplocos sumuntia	04	U	E	乌饭树 Vaccinium bracteatum	44	S	E	麂角杜鹃 Rhododendron latoucheae	46	U	E
香冬青 Ilex suaveolens	03	T	E	木荷 Schima superba	43	T	E	短尾越桔Vaccinium carlesii	44	S	E

	群丛1 Asso. 1	群丛2 Asso. 2	群丛3 Asso. 3
面积 Area (hm²)	12.4	3.32	8.28
丰富度 Richness	145	108	145
个体密度 Individual density (stem· hm^-2)	5 260	9 961	5 121
平均胸径 Mean DBH (cm)	5.73	3.95	5.32