Chin J Plant Ecol ›› 2010, Vol. 34 ›› Issue (4): 409-417.DOI: 10.3773/j.issn.1005-264x.2010.04.006
• Research Articles • Previous Articles Next Articles
SUN Hong-Gang1, ZHANG Jian-Guo2,*(), DUAN Ai-Guo2
Received:
2009-06-09
Accepted:
2010-01-25
Online:
2010-06-09
Published:
2010-04-01
Contact:
ZHANG Jian-Guo
SUN Hong-Gang, ZHANG Jian-Guo, DUAN Ai-Guo. A comparison of selecting data points and fitting coefficients methods for estimating self-thinning boundary line[J]. Chin J Plant Ecol, 2010, 34(4): 409-417.
样地 Plot | 初植密度 Planting density (trees·hm-2) | 2006年林分密度 Stand density in 2006 (trees·hm-2) | 林分平均胸径 Mean diameter of breast height (cm) | 标准差 SD | 胸径最小值 Minimum diameter of breast height (cm) | 胸径最大值 Maximum diameter of breast height (cm) |
---|---|---|---|---|---|---|
B2 | 3 333 | 2 517 | 10.65 | 2.42 | 6.49 | 13.80 |
B3 | 3 333 | 2 483 | 12.29 | 2.71 | 7.58 | 16.23 |
C2 | 5 000 | 3 467 | 9.57 | 2.44 | 5.51 | 13.16 |
C3 | 5 000 | 2 417 | 10.21 | 2.74 | 6.04 | 14.04 |
D1 | 6 667 | 3 317 | 8.76 | 2.02 | 5.53 | 11.90 |
D2 | 6 667 | 4 267 | 8.85 | 2.67 | 4.95 | 13.14 |
D3 | 6 667 | 3 800 | 9.13 | 2.62 | 5.33 | 13.26 |
E1 | 10 000 | 5 117 | 8.39 | 2.01 | 5.55 | 11.70 |
E2 | 10 000 | 3 183 | 8.42 | 2.50 | 4.80 | 13.08 |
E3 | 10 000 | 5 217 | 8.75 | 2.71 | 5.10 | 13.11 |
Table 1 Descriptive statistics of plots
样地 Plot | 初植密度 Planting density (trees·hm-2) | 2006年林分密度 Stand density in 2006 (trees·hm-2) | 林分平均胸径 Mean diameter of breast height (cm) | 标准差 SD | 胸径最小值 Minimum diameter of breast height (cm) | 胸径最大值 Maximum diameter of breast height (cm) |
---|---|---|---|---|---|---|
B2 | 3 333 | 2 517 | 10.65 | 2.42 | 6.49 | 13.80 |
B3 | 3 333 | 2 483 | 12.29 | 2.71 | 7.58 | 16.23 |
C2 | 5 000 | 3 467 | 9.57 | 2.44 | 5.51 | 13.16 |
C3 | 5 000 | 2 417 | 10.21 | 2.74 | 6.04 | 14.04 |
D1 | 6 667 | 3 317 | 8.76 | 2.02 | 5.53 | 11.90 |
D2 | 6 667 | 4 267 | 8.85 | 2.67 | 4.95 | 13.14 |
D3 | 6 667 | 3 800 | 9.13 | 2.62 | 5.33 | 13.26 |
E1 | 10 000 | 5 117 | 8.39 | 2.01 | 5.55 | 11.70 |
E2 | 10 000 | 3 183 | 8.42 | 2.50 | 4.80 | 13.08 |
E3 | 10 000 | 5 217 | 8.75 | 2.71 | 5.10 | 13.11 |
Fig. 1 Relationship between stand density (N) and mean diameter of breast height (D) under four regression methods. A, Visualized inspection. B, Mortality criterion. C, Equal intervals. D, Relative density (RD).
数据点选择方法及数量 Selecting data points and numbers (n) | 斜率(标准误差) Slope (SE) | 截距(标准误差) Intercept (SE) | R2 |
---|---|---|---|
死亡率法 Mortality criterion | |||
开始死亡 Zero mortality (83) | -0.56 (0.046) | 7.28 (0.612) | 0.817 |
10%死亡 10% mortality (51) | -0.51 (0.037) | 6.79 (0.435) | 0.931 |
20%死亡 20% mortality (37) | -0.49 (0.029) | 6.19 (0.367) | 0.934 |
等距区间法 Equal intervals | |||
5株 5 stems (43) | -0.62 (0.071) | 7.91 (0.610) | 0.917 |
10株 10 stems (31) | -0.55 (0.069) | 7.26 (0.583) | 0.922 |
20株 20 stems (19) | -0.54 (0.089) | 7.23 (0.762) | 0.939 |
相对密度法 Relative density (RD) | |||
RD > 0.7 (71) | -0.58 (0.069) | 7.14 (0.583) | 0.883 |
RD > 0.8 (62) | -0.54 (0.056) | 7.09 (0.472) | 0.929 |
RD > 0.9 (57) | -0.53 (0.057) | 6.70 (0.478) | 0.900 |
Table 2 Comparison of the selecting data points and fitting coefficients
数据点选择方法及数量 Selecting data points and numbers (n) | 斜率(标准误差) Slope (SE) | 截距(标准误差) Intercept (SE) | R2 |
---|---|---|---|
死亡率法 Mortality criterion | |||
开始死亡 Zero mortality (83) | -0.56 (0.046) | 7.28 (0.612) | 0.817 |
10%死亡 10% mortality (51) | -0.51 (0.037) | 6.79 (0.435) | 0.931 |
20%死亡 20% mortality (37) | -0.49 (0.029) | 6.19 (0.367) | 0.934 |
等距区间法 Equal intervals | |||
5株 5 stems (43) | -0.62 (0.071) | 7.91 (0.610) | 0.917 |
10株 10 stems (31) | -0.55 (0.069) | 7.26 (0.583) | 0.922 |
20株 20 stems (19) | -0.54 (0.089) | 7.23 (0.762) | 0.939 |
相对密度法 Relative density (RD) | |||
RD > 0.7 (71) | -0.58 (0.069) | 7.14 (0.583) | 0.883 |
RD > 0.8 (62) | -0.54 (0.056) | 7.09 (0.472) | 0.929 |
RD > 0.9 (57) | -0.53 (0.057) | 6.70 (0.478) | 0.900 |
回归方法 Regression technique | 斜率 Slope | 截距 Intercept |
---|---|---|
随机边界方程 Stochastic frontier function | -0.52 | 7.03 |
最小二乘法 Ordinary least squares regression | -0.54 | 7.63 |
降维分析法 Reduced major axis | -0.57 | 7.39 |
分位数回归法 Quantile regression | -0.52 | 7.05 |
Table 3 Regression coefficients of the four models
回归方法 Regression technique | 斜率 Slope | 截距 Intercept |
---|---|---|
随机边界方程 Stochastic frontier function | -0.52 | 7.03 |
最小二乘法 Ordinary least squares regression | -0.54 | 7.63 |
降维分析法 Reduced major axis | -0.57 | 7.39 |
分位数回归法 Quantile regression | -0.52 | 7.05 |
[1] | Bi H (2001). The self-thinning surface. Forest Science, 47, 361-370. |
[2] | Bi H, Turvey ND (1997). A method of selecting data points for fitting the maximum biomass-density line for stand undergoing self-thinning. Australian Journal of Ecology, 22, 356-359. |
[3] | Bi H, Wan G, Turvey ND (2000). Estimating the self-thinning boundary line as a density-dependent stochastic biomass frontier. Ecology, 81, 1477-1483. |
[4] | Cade BS, Terrell JW, Schroeder RL (1999). Estimating effects of limiting factors with regression quantiles. Ecology, 80, 311-323. |
[5] | Drew TJ, Flewelling JW (1979). Stand density management: an alternative approach and its application to Douglas-fir plantations. Forest Science, 25, 518-532. |
[6] | Econometric Software (1998). LIMDEP 7. 0 User’s Manual (Computer Manual). Econometric Software. Plainview, New York, USA. |
[7] | Enquist BJ, Brown JH, West GB (1998) . Allometric scaling of plant energetics and population density. Nature, 395, 163-165. |
[8] | Fang JY (方精云) (1992). Self-thinning rule in plant population. Rural Eco-Environment (农村生态环境), 2, 7-12. (in Chinese with English abstract) |
[9] | Fang JY (方精云), Jian C (菅诚), Yamakura T (1991). Relationships between population growth and population density in monocultures of Larix leptolepis. Acta Botanica Sinica (植物学报), 33, 949-957. (in Chinese with English abstract) |
[10] | Gorham E (1979). Shoot height, weight and standing crop in relation to density of monospecific plant stands. Nature, 279, 148-150. |
[11] | Jack SB, Long JN (1996). Linkages between silviculture and ecology: an analysis of density management diagrams. Forest Ecology and Management, 86, 205-220. |
[12] | Kikuzawa K (1999). Theoretical relationships between mean plant size, size distribution and self thinning under one-sided competition. Annals of Botany, 83, 11-18. |
[13] | Koenker R, Bassett G (1978). Regression quantiles. Econometrica, 46, 33-50. |
[14] | LaBarbera M (1989). Analyzing body size as factor in ecology and evolution. Annual Review of Ecology and Systematics, 20, 97-117. |
[15] | Li B, Wu H, Zou G (2000). Self-thinning rule: a causal interpretation from ecological field theory. Ecological Modelling, 132, 167-173. |
[16] | Lonsdale W (1990). The self-thinning rule: dead or alive? Ecology, 71, 1373-1388. |
[17] |
Long JN, Smith FW (1984). Relation between size and density in developing stands: a description and possible mechanisms. Forest Ecology and Management, 7, 191-206.
DOI URL |
[18] |
Mohler C, Marks P, Sprugel DG (1978). Stand structure and allometry of trees during self-thinning of pure stands. Journal of Ecology, 66, 599-614.
DOI URL |
[19] | Newton PF (2006). Asymptotic size-density relationships within self-thinning black spruce and jack pine stand- types: parameter estimation and model reformulations. Forest Ecology and Management, 226, 49-59. |
[20] | Osawa A (1995). Inverse relationship of crown fractal dimension to self-thinning exponent of tree population: a hypothesis. Canadian Journal of Forestry Research, 25, 1608-1617. |
[21] |
Osawa A, Allen RB (1993). Allometric theory explains self-thinning relationships of mountain beech and red pine. Ecology, 74, 1020-1032.
DOI URL |
[22] |
Osawa A, Sugita S (1989). The self-thinning rule: another interpretation of Weller’s results. Ecology, 70, 279-283.
DOI URL |
[23] | Río M, Montero G, Bravo F (2001). Analysis of diameter- density relationships and self-thinning in non-thinned even-aged Scots pine stands. Forest Ecology and Management, 142, 79-87. |
[24] | Roderick ML, Barnes B (2004). Self-thinning of plant populations from a dynamic viewpoint. Functional Ecology, 18, 197-203. |
[25] | SAS Institute (1999). SAS/STAT User's Guide, Version 8 (Computer Manual). SAS Institute Inc., Cary, New York, USA. |
[26] | Scharf FS, Juanes F, Sutherland M (1998). Inferring ecological relationships from the edges of scatter diagrams: comparison of regression techniques. Ecology, 79, 448-460. |
[27] | Solomon DS, Zhang L (2002). Maximum size-density relationships for mixed softwoods in the northeastern USA. Forest Ecology and Management, 155, 163-170. |
[28] | Weller DE (1987). A reevaluation of the -3/2 power rule of plant self-thinning. Ecological Monographs, 57, 23-43. |
[29] | Weller DE (1989). The interspecific size-density relationship among crowded plant stands and its implications for the -3/2 power rule of self-thinning. American Naturalist, 133, 20-41. |
[30] | Westoby M (1984). The self-thinning rule. Advances in Ecological Research, 14, 167-225. |
[31] | White J (1981). The allometric interpretation of the self- thinning rule. Journal of Theoretical Biology, 89, 475-500. |
[32] | Wilson DS, Seymour RS, Maguire DA (1999). Density management diagram for Northeastern red spruce and balsam fir forests. Northern Journal of Applied Forestry, 16, 48-56. |
[33] | Yang Y, Titus SJ (2002). Maximum size-density relationship for constraining individual tree mortality functions. Forest Ecology and Management, 168, 259-273. |
[34] | Yoda K, Kira T, Ogawa H (1963). Self-thinning in overcrowded pure stand under cultivated and natural conditions. Journal of Biology of Osaka City University, 14, 107-129. |
[35] | Zeide B (1987). Analysis of the 3/2 power law of self-thinning. Forest Science, 33, 517-537. |
[1] | XIE Huan, ZHANG Qiu-Fang, ZENG Quan-Xin, ZHOU Jia-Cong, MA Ya-Pei, WU Yue, LIU Yuan-Yuan, LIN Hui-Ying, YIN Yun-Feng, CHEN Yue-Min. Effects of nitrogen addition on phosphorus transformation and decomposition fungi in seedling stage of Cunninghamia lanceolata [J]. Chin J Plant Ecol, 2022, 46(2): 220-231. |
[2] | LÜ Zhong-Cheng, KANG Wen-Xing, HUANG Zhi-Hong, ZHAO Zhong-Hui, DENG Xiang-Wen. Reuse of retranslocated nutrients in tissues of Chinese fir in plantations of different ages [J]. Chin J Plant Ecol, 2019, 43(5): 458-470. |
[3] | SHEN Fang-Fang, LI Yan-Yan, LIU Wen-Fei, DUAN Hong-Lang, FAN Hou-Bao, HU Liang, MENG Qing-Yin. Responses of nitrogen and phosphorus resorption from leaves and branches to long-term nitrogen deposition in a Chinese fir plantation [J]. Chin J Plant Ecol, 2018, 42(9): 926-937. |
[4] | PENG Xi, YAN Wen-De, WANG Feng-Qi, WANG Guang-Jun, YU Fang-Yong, ZHAO Mei-Fang. Specific leaf area estimation model building based on leaf dry matter content of Cunninghamia lanceolata [J]. Chin J Plant Ecol, 2018, 42(2): 209-219. |
[5] | CHEN Ri-Sheng, KANG Wen-Xing, ZHOU Yu-Quan, TIAN Da-Lun, XIANG Wen-Hua . Changes in nutrient cycling with age in a Cunninghamia lanceolata plantation forest [J]. Chin J Plan Ecolo, 2018, 42(2): 173-184. |
[6] | CHEN Si-Tong, ZOU Xian-Hua, CAI Yi-Bing, WEI Dan, LI Tao, WU Peng-Fei, MA Xiang-Qing. Phosphorus distribution inside Chinese fir seedlings under different P supplies based on 32P tracer [J]. Chin J Plant Ecol, 2018, 42(11): 1103-1112. |
[7] | Shun-Zeng SHI, De-Cheng XIONG, Fei DENG, Jian-Xin FENG, Chen-Sen XU, Bo-Yuan ZHONG, Yun-Yu CHEN, Guang-Shui CHEN, Yu-Sheng YANG. Interactive effects of soil warming and nitrogen addition on fine root production of Chinese fir seedlings [J]. Chin J Plan Ecolo, 2017, 41(2): 186-195. |
[8] | Zhi-Yu CHEN, Qi LI, Xian-Hua ZOU, Xiang-Qing MA, Peng-Fei WU. Effect of neighboring competition on photosynthetic characteristics and biomass allocation of Chinese fir seedlings under low phosphorus stress [J]. Chin J Plant Ecol, 2016, 40(2): 177-. |
[9] | Xiang GU, Shi-Ji ZHANG, Wen-Hua XIANG, Lei-Da LI, Zhao-Dan LIU, Wei-Jun SUN, Xi FANG. Seasonal dynamics of active soil organic carbon in four subtropical forests in Southern China [J]. Chin J Plant Ecol, 2016, 40(10): 1064-1076. |
[10] | WAN Jing-Juan,GUO Jian-Fen,JI Shu-Rong,REN Wei-Ling,SI You-Tao,YANG Yu-Sheng. Effects of different sources of dissolved organic matter on soil CO2 emission in subtropical forests [J]. Chin J Plan Ecolo, 2015, 39(7): 674-681. |
[11] | WANG Qing-Kui,LI Yan-Peng,ZHANG Fang-Yue,HE Tong-Xin. Short-term nitrogen fertilization decreased root and microbial respiration in a young Cunninghamia lanceolata plantation [J]. Chin J Plan Ecolo, 2015, 39(12): 1166-1175. |
[12] | LIU Rong-Jie, WU Ya-Cong, ZHANG Ying, LI Zheng-Cai, MA Shao-Jie, WANG Bin, GERI Le-Tu. Comparison of soil labile organic carbon in Chinese fir plantations and natural secondary forests in north subtropical areas of China [J]. Chin J Plant Ecol, 2012, 36(5): 431-437. |
[13] | LIU Jia, XIANG Wen-Hua, XU Xiao, CHEN Rui, TIAN Da-Lun, PENG Chang-Hui, FANG Xi. Analysis of architecture and functions of fine roots of five subtropical tree species in Huitong, Hunan Province, China [J]. Chin J Plant Ecol, 2010, 34(8): 938-945. |
[14] | HE Qian, LI Ji-Yue, CHEN Xiao-Yang, CHEN Hong-Yue, PENG Hua-Gui, FAN Shun-Jiang. Types and extent of damage to Cunninghamia lanceolata plantations due to unusually heavy snow and ice in southern China [J]. Chin J Plant Ecol, 2010, 34(2): 195-203. |
[15] | WANG Guang-Jun, TIAN Da-Lun, YAN Wen-De, ZHU Fan, XIANG Wen-Hua, LIANG Xiao-Cui. EFFECTS OF ABOVEGROUND LITTER EXCLUSION AND ADDITION ON SOIL RESPIRATION IN A CUNNINGHAMIA LANCEOLATA PLANTATION IN CHINA [J]. Chin J Plant Ecol, 2009, 33(4): 739-747. |
Viewed | ||||||
Full text |
|
|||||
Abstract |
|
|||||
Copyright © 2022 Chinese Journal of Plant Ecology
Tel: 010-62836134, 62836138, E-mail: apes@ibcas.ac.cn, cjpe@ibcas.ac.cn