植物生态学报 ›› 2013, Vol. 37 ›› Issue (4): 306-316.DOI: 10.3724/SP.J.1258.2013.00030
收稿日期:
2012-10-22
接受日期:
2013-02-25
出版日期:
2013-10-22
发布日期:
2013-04-09
通讯作者:
杨万勤
作者简介:
*(E-mail:scyangwq@163.com)基金资助:
HE Wei, WU Fu-Zhong, YANG Wan-Qin*(), WU Qi-Qian, HE Min, ZHAO Ye-Yi
Received:
2012-10-22
Accepted:
2013-02-25
Online:
2013-10-22
Published:
2013-04-09
Contact:
YANG Wan-Qin
摘要:
高山/亚高山森林灌木层植物凋落物的分解对于系统物质循环等过程具有重要意义, 并可能受到冬季不同厚度雪被斑块下冻融格局的影响。该文采用凋落物分解袋法, 研究了高山森林典型灌层植物华西箭竹(Fargesia nitida)和康定柳(Salix paraplesia)凋落物在沿林窗-林下形成的冬季雪被厚度梯度(厚型雪被斑块、较厚型雪被斑块、中型雪被斑块、薄型雪被斑块、无雪被斑块)上在第一年不同关键时期(冻结初期、冻结期、融化期、生长季节初期和生长季节后期)的质量损失特征。在整个冻融季节, 华西箭竹和康定柳凋落叶的平均质量损失分别占全年的(48.78 ± 2.35)%和(46.60 ± 5.02)%。冻融季节雪被覆盖斑块下凋落叶的失重率表现出厚型雪被斑块大于薄型雪被斑块的趋势,而生长季节无雪被斑块的失重率明显较高。尽管如此, 华西箭竹凋落物第一年分解表现出随冬季雪被厚度增加而增加的趋势, 但康定柳凋落物第一年失重率以薄型雪被斑块最高, 而无雪被斑块最低。同时, 相关分析表明冻融季节凋落叶的失重率与平均温度和负积温呈极显著正相关, 生长季节凋落叶的失重率与所调查的温度因子并无显著相关关系, 但全年凋落物失重率与平均温度和正/负积温均显著相关。这些结果清晰地表明, 未来冬季变暖情境下高山森林冬季雪被格局的改变将显著影响灌层植物凋落物分解, 影响趋势随着物种的差异具有明显差异。
何伟, 吴福忠, 杨万勤, 武启骞, 何敏, 赵野逸. 雪被斑块对高山森林两种灌木凋落叶质量损失的影响. 植物生态学报, 2013, 37(4): 306-316. DOI: 10.3724/SP.J.1258.2013.00030
HE Wei, WU Fu-Zhong, YANG Wan-Qin, WU Qi-Qian, HE Min, ZHAO Ye-Yi. Effect of snow patches on leaf litter mass loss of two shrubs in an alpine forest. Chinese Journal of Plant Ecology, 2013, 37(4): 306-316. DOI: 10.3724/SP.J.1258.2013.00030
物种 Species | C (g·kg-1) | N (g·kg-1) | P (g·kg-1) | C/N | C/P | N/P | 木质素 Lignin (%) | 纤维素 Cellulose (%) | 木质素/ 纤维素 Lignin/ Cellulose | 木质素/氮 Lignin/N |
---|---|---|---|---|---|---|---|---|---|---|
华西箭竹 Fargesia nitida | 317.71 ± 16.60a | 9.02 ± 0.12b | 0.94 ± 0.07b | 35.23 ± 1.38a | 339.80 ± 9.11a | 9.66 ± 0.64b | 14.79 ± 0.62b | 12.97 ± 0.48a | 1.14 ± 0.00b | 16.40 ± 0.47a |
康定柳 Salix paraplesia | 371.89 ± 31.55a | 14.33 ± 0.26a | 1.28 ± 0.06a | 25.93 ± 1.74b | 290.72 ± 10.31b | 11.23 ± 0.36a | 21.79 ± 1.02a | 10.60 ± 1.04b | 2.06 ± 1.11a | 15.20 ± 0.44b |
表1 华西箭竹和康定柳凋落叶初始质量(平均值±标准偏差, n = 5)
Table 1 Initial quality in leaf litters of Fargesia nitida and Salix paraplesia (mean ± SD, n = 5)
物种 Species | C (g·kg-1) | N (g·kg-1) | P (g·kg-1) | C/N | C/P | N/P | 木质素 Lignin (%) | 纤维素 Cellulose (%) | 木质素/ 纤维素 Lignin/ Cellulose | 木质素/氮 Lignin/N |
---|---|---|---|---|---|---|---|---|---|---|
华西箭竹 Fargesia nitida | 317.71 ± 16.60a | 9.02 ± 0.12b | 0.94 ± 0.07b | 35.23 ± 1.38a | 339.80 ± 9.11a | 9.66 ± 0.64b | 14.79 ± 0.62b | 12.97 ± 0.48a | 1.14 ± 0.00b | 16.40 ± 0.47a |
康定柳 Salix paraplesia | 371.89 ± 31.55a | 14.33 ± 0.26a | 1.28 ± 0.06a | 25.93 ± 1.74b | 290.72 ± 10.31b | 11.23 ± 0.36a | 21.79 ± 1.02a | 10.60 ± 1.04b | 2.06 ± 1.11a | 15.20 ± 0.44b |
图1 川西高山森林不同雪被斑块的雪被覆盖厚度(平均值±标准偏差, n = 5)。 SP1, 厚型雪被斑块; SP2, 较厚型雪被斑块; SP3, 中型雪被斑块; SP4, 薄型雪被斑块。
Fig. 1 Thickness of snow cover under different snow patches in alpine forest of western Sichuan (mean ± SD, n = 5). SP1, thickest snow cover patch; SP2, thicker snow cover patch; SP3, middle thick snow cover patch; SP4, thinner snow cover patch.
土壤冻结 初期 Onset of soil freezing period | 土壤冻 结期 Soil freezing period | 土壤融化期 Soil thawing period | 生长季节初期 Early growth period | 生长季节 后期 Later growth period | 冻融季节 Freeze- thaw season | 生长季节 Growth season | 全年 Whole year | ||
---|---|---|---|---|---|---|---|---|---|
厚型雪被斑块 Thickest snow cover patch | 平均温度 AT | 0.38 | -2.00 | -0.14 | 12.33 | 6.10 | -0.75 | 8.77 | 3.60 |
正积温 PAT | 35.35 | – | 1.08 | 1501.94 | 489.21 | 36.43 | 1 991.15 | 2 027.58 | |
负积温 NAT | -19.14 | -139.23 | -7.79 | – | – | -166.16 | – | -166.16 | |
冻融循环次数 FSFC | 48.00 | – | 2.00 | 15.00 | – | 50.00 | 15.00 | 65.00 | |
较厚型雪被斑块 Thicker snow cover patch | 平均温度 AT | -0.49 | -2.40 | 1.00 | 10.80 | 6.74 | -0.86 | 8.48 | 3.39 |
正积温 PAT | 19.19 | 6.70 | 49.97 | 1311.93 | 543.60 | 75.86 | 1 855.52 | 1 931.38 | |
负积温 NAT | -45.31 | -173.65 | -2.82 | – | – | -221.79 | – | -221.79 | |
冻融循环次数 FSFC | 49.00 | 54.00 | 23.00 | 13.00 | – | 126.00 | 13.00 | 139.00 | |
中型雪被斑块 Middle thick snow cover patch | 平均温度 AT | -0.66 | -2.86 | 0.37 | 7.86 | 6.08 | -1.28 | 6.84 | 2.43 |
正积温 PAT | 10.87 | – | 29.64 | 960.82 | 487.88 | 40.50 | 1 448.70 | 1 489.20 | |
负积温 NAT | -45.47 | -199.81 | -12.03 | – | – | -257.31 | – | -257.31 | |
冻融循环次数 FSFC | 41.00 | 5.00 | 18.00 | 13.00 | – | 64.00 | 13.00 | 77.00 | |
薄型雪被斑块 Thinner snow cover patch | 平均温度 AT | -0.61 | -3.53 | 0.32 | 6.90 | 5.43 | -1.56 | 6.06 | 1.92 |
正积温 PAT | 22.73 | 0.61 | 38.22 | 844.17 | 434.61 | 61.56 | 1 278.78 | 1 340.35 | |
负积温 NAT | -55.36 | -246.35 | -22.73 | – | – | -324.44 | – | -324.44 | |
冻融循环次数 FSFC | 38.00 | 22.00 | 34.00 | 1.00 | – | 94.00 | 1.00 | 95.00 | |
无雪被斑块 No snow cover patch | 平均温度 AT | -1.02 | -3.47 | 0.50 | 7.67 | 5.19 | -1.61 | 6.68 | 1.50 |
正积温 PAT | 10.60 | – | 41.56 | 938.53 | 421.46 | 52.16 | 1 359.99 | 1 412.15 | |
负积温 NAT | -64.84 | -241.50 | -17.49 | – | -6.76 | -323.82 | -6.76 | -330.58 | |
冻融循环次数 FSFC | 39.00 | 15.00 | 38.00 | – | 27.00 | 92.00 | 27.00 | 119.00 |
表2 不同雪被斑块各分解时期的土壤平均温度、正积温、负积温和土壤冻融循环次数特征
Table 2 Soil average temperature (AT), positive accumulated temperature (PAT), negative accumulated temperature (NAT) and frequency of soil freeze-thaw cycle (FSFC) in each decomposition stage under different snow patches
土壤冻结 初期 Onset of soil freezing period | 土壤冻 结期 Soil freezing period | 土壤融化期 Soil thawing period | 生长季节初期 Early growth period | 生长季节 后期 Later growth period | 冻融季节 Freeze- thaw season | 生长季节 Growth season | 全年 Whole year | ||
---|---|---|---|---|---|---|---|---|---|
厚型雪被斑块 Thickest snow cover patch | 平均温度 AT | 0.38 | -2.00 | -0.14 | 12.33 | 6.10 | -0.75 | 8.77 | 3.60 |
正积温 PAT | 35.35 | – | 1.08 | 1501.94 | 489.21 | 36.43 | 1 991.15 | 2 027.58 | |
负积温 NAT | -19.14 | -139.23 | -7.79 | – | – | -166.16 | – | -166.16 | |
冻融循环次数 FSFC | 48.00 | – | 2.00 | 15.00 | – | 50.00 | 15.00 | 65.00 | |
较厚型雪被斑块 Thicker snow cover patch | 平均温度 AT | -0.49 | -2.40 | 1.00 | 10.80 | 6.74 | -0.86 | 8.48 | 3.39 |
正积温 PAT | 19.19 | 6.70 | 49.97 | 1311.93 | 543.60 | 75.86 | 1 855.52 | 1 931.38 | |
负积温 NAT | -45.31 | -173.65 | -2.82 | – | – | -221.79 | – | -221.79 | |
冻融循环次数 FSFC | 49.00 | 54.00 | 23.00 | 13.00 | – | 126.00 | 13.00 | 139.00 | |
中型雪被斑块 Middle thick snow cover patch | 平均温度 AT | -0.66 | -2.86 | 0.37 | 7.86 | 6.08 | -1.28 | 6.84 | 2.43 |
正积温 PAT | 10.87 | – | 29.64 | 960.82 | 487.88 | 40.50 | 1 448.70 | 1 489.20 | |
负积温 NAT | -45.47 | -199.81 | -12.03 | – | – | -257.31 | – | -257.31 | |
冻融循环次数 FSFC | 41.00 | 5.00 | 18.00 | 13.00 | – | 64.00 | 13.00 | 77.00 | |
薄型雪被斑块 Thinner snow cover patch | 平均温度 AT | -0.61 | -3.53 | 0.32 | 6.90 | 5.43 | -1.56 | 6.06 | 1.92 |
正积温 PAT | 22.73 | 0.61 | 38.22 | 844.17 | 434.61 | 61.56 | 1 278.78 | 1 340.35 | |
负积温 NAT | -55.36 | -246.35 | -22.73 | – | – | -324.44 | – | -324.44 | |
冻融循环次数 FSFC | 38.00 | 22.00 | 34.00 | 1.00 | – | 94.00 | 1.00 | 95.00 | |
无雪被斑块 No snow cover patch | 平均温度 AT | -1.02 | -3.47 | 0.50 | 7.67 | 5.19 | -1.61 | 6.68 | 1.50 |
正积温 PAT | 10.60 | – | 41.56 | 938.53 | 421.46 | 52.16 | 1 359.99 | 1 412.15 | |
负积温 NAT | -64.84 | -241.50 | -17.49 | – | -6.76 | -323.82 | -6.76 | -330.58 | |
冻融循环次数 FSFC | 39.00 | 15.00 | 38.00 | – | 27.00 | 92.00 | 27.00 | 119.00 |
图2 川西高山森林不同雪被斑块下土壤表层和大气的昼平均温度(7:00-19:00)与夜平均温度(19:00-7:00)动态(2010年10月26日至2011年11月8日)。 SP1, 厚型雪被斑块; SP2, 较厚型雪被斑块; SP3, 中型雪被斑块; SP4, 薄型雪被斑块; SP5, 无雪被斑块。
Fig. 2 Dynamics of average temperature at daytime (7:00-19:00) and nighttime (19:00-7:00) between surface soil and atmosphere under different snow patches in alpine forest of western Sichuan from 26 October 2010 to 8 November 2011. SP1, thickest snow cover patch; SP2, thicker snow cover patch; SP3, middle thick snow cover patch; SP4, thinner snow cover patch; SP5, no snow cover patch.
图3 川西高山森林不同雪被斑块下华西箭竹和康定柳凋落叶的失重率(2010年10月26日至2011年11月8日) (平均值±标准偏差, n = 3)。 不同小写字母表示相同分解期不同斑块间差异显著(p < 0.05)。SP1, 厚型雪被斑块; SP2, 较厚型雪被斑块; SP3, 中型雪被斑块; SP4, 薄型雪被斑块; SP5, 无雪被斑块。
Fig. 3 Mass loss rates of Fargesia nitida and Salix paraplesia leaf litter under different snow patches in alpine forest of western Sichuan from 26 October 2010 to 8 November 2011 (mean ± SD, n = 3). Different lowercase letters indicate significant difference (p < 0.05) among different snow patches within the same decomposition stage. SP1, thickest snow cover patch; SP2, thicker snow cover patch; SP3, middle thick snow cover patch; SP4, thinner snow cover patch; SP5, no snow cover patch.
图4 川西高山森林不同斑块下华西箭竹和康定柳凋落叶各分解阶段对全年质量损失的贡献率。 SP1, 厚型雪被斑块; SP2, 较厚型雪被斑块; SP3, 中型雪被斑块; SP4, 薄型雪被斑块; SP5, 无雪被斑块。括号内数字表示距前次采样间隔天数。
Fig. 4 Contribution rates (%) of Fargesia nitida and Salix paraplesia leaf litter at each decomposition stage to one-year mass lose under different snow patches in alpine forest of western Sichuan. SP1, thickest snow cover patch; SP2, thicker snow cover patch; SP3, middle thick snow cover patch; SP4, thinner snow cover patch; SP5, no snow cover patch. Numbers in brackets indicate the days between corresponding and former sampling date.
物种 Species | 雪被斑块 Snow patches | 23 Dec. 2010 (58) | 3 Mar. 2011 (70) | 19 Apr. 2011 (47) | 19 Aug. 2011 (122) | 8 Nov. 2011 (81) |
---|---|---|---|---|---|---|
华西箭竹 | SP1 | 3.80 ± 0.16b | 3.77 ± 0.37b | 1.67 ± 0.18c | 4.83 ± 0.25a | 3.91 ± 0.18b |
Fargesia nitida | SP2 | 4.24 ± 0.11ab | 2.74 ± 0.47c | 1.88 ± 0.52d | 4.60 ± 0.46a | 3.97 ± 0.41b |
SP3 | 3.63 ± 0.17d | 2.00 ± 0.18c | 1.86 ± 0.34c | 4.62 ± 0.30a | 4.06 ± 0.37b | |
SP4 | 4.69 ± 0.28a | 1.78 ± 0.16c | 1.59 ± 0.36c | 4.83 ± 0.13a | 3.62 ± 0.26b | |
SP5 | 4.20 ± 0.12b | 1.74 ± 0.23d | 1.47 ± 0.20d | 5.38 ± 0.44a | 3.15 ± 0.33c | |
康定柳 | SP1 | 2.86 ± 0.28d | 6.77 ± 0.37a | 1.74 ± 0.40e | 5.11 ± 0.51b | 4.47 ± 0.22c |
Salix paraplesia | SP2 | 2.28 ± 0.26c | 5.93 ± 0.24a | 1.75 ± 0.41d | 5.50 ± 0.33b | 5.71 ± 0.15ab |
SP3 | 2.71 ± 0.14b | 4.78 ± 0.61a | 0.90 ± 0.08c | 4.72 ± 0.12a | 4.89 ± 0.25a | |
SP4 | 2.15 ± 0.22c | 4.40 ± 0.20b | 1.37 ± 0.40d | 5.45 ± 0.26a | 4.37 ± 0.38b | |
SP5 | 2.28 ± 0.16d | 3.67 ± 0.36c | 0.84 ± 0.13e | 5.53 ± 0.25a | 4.47 ± 0.42b |
表3 川西高山森林不同雪被斑块下华西箭竹凋落叶和康定柳凋落叶每30天的失重速率(2010年10月26日至2011年11月8日) (平均值±标准误差, n = 3)
Table 3 Mass loss rate per 30 days of Fargesia nitida and Salix paraplesia leaf litter under different snow patches in alpine forest of western Sichuan from 26 October 2010 to 8 November 2011 (mean ± SE, n = 3)
物种 Species | 雪被斑块 Snow patches | 23 Dec. 2010 (58) | 3 Mar. 2011 (70) | 19 Apr. 2011 (47) | 19 Aug. 2011 (122) | 8 Nov. 2011 (81) |
---|---|---|---|---|---|---|
华西箭竹 | SP1 | 3.80 ± 0.16b | 3.77 ± 0.37b | 1.67 ± 0.18c | 4.83 ± 0.25a | 3.91 ± 0.18b |
Fargesia nitida | SP2 | 4.24 ± 0.11ab | 2.74 ± 0.47c | 1.88 ± 0.52d | 4.60 ± 0.46a | 3.97 ± 0.41b |
SP3 | 3.63 ± 0.17d | 2.00 ± 0.18c | 1.86 ± 0.34c | 4.62 ± 0.30a | 4.06 ± 0.37b | |
SP4 | 4.69 ± 0.28a | 1.78 ± 0.16c | 1.59 ± 0.36c | 4.83 ± 0.13a | 3.62 ± 0.26b | |
SP5 | 4.20 ± 0.12b | 1.74 ± 0.23d | 1.47 ± 0.20d | 5.38 ± 0.44a | 3.15 ± 0.33c | |
康定柳 | SP1 | 2.86 ± 0.28d | 6.77 ± 0.37a | 1.74 ± 0.40e | 5.11 ± 0.51b | 4.47 ± 0.22c |
Salix paraplesia | SP2 | 2.28 ± 0.26c | 5.93 ± 0.24a | 1.75 ± 0.41d | 5.50 ± 0.33b | 5.71 ± 0.15ab |
SP3 | 2.71 ± 0.14b | 4.78 ± 0.61a | 0.90 ± 0.08c | 4.72 ± 0.12a | 4.89 ± 0.25a | |
SP4 | 2.15 ± 0.22c | 4.40 ± 0.20b | 1.37 ± 0.40d | 5.45 ± 0.26a | 4.37 ± 0.38b | |
SP5 | 2.28 ± 0.16d | 3.67 ± 0.36c | 0.84 ± 0.13e | 5.53 ± 0.25a | 4.47 ± 0.42b |
不同分解阶段失重率 Mass lose rates at different decomposition stages | 土壤平均温度 Soil average temperature | 正积温 Positive accumulated temperature | 负积温 Negative accumulated temperature | 冻融循环次数 Frequency of soil freeze-thaw cycle |
---|---|---|---|---|
土壤冻结初期 Onset of soil freezing period | 0.031 | 0.060 | 0.010 | -0.009 |
土壤冻结期 Soil freezing period | 0.533** | 0.157 | 0.534** | -0.008 |
土壤融化期 Soil thawing period | 0.061 | -0.100 | 0.378* | -0.319 |
生长季节初期 Early growth period | -0.117 | -0.117 | – | -0.431* |
生长季节后期 Later growth period | 0.486** | 0.487** | 0.313 | -0.313 |
冻融季节 Freeze-thaw season | 0.751** | -0.051 | 0.733** | -0.148 |
生长季节 Growth season | 0.151 | 0.133 | 0.027 | 0.029 |
全年 Whole year | 0.589** | 0.582** | 0.547** | -0.023 |
Table 4 Correlation analyses among mass lose rate (%), soil average temperature, positive accumulated temperature, negative accumulated temperature, and frequency of soil freeze-thaw cycle
不同分解阶段失重率 Mass lose rates at different decomposition stages | 土壤平均温度 Soil average temperature | 正积温 Positive accumulated temperature | 负积温 Negative accumulated temperature | 冻融循环次数 Frequency of soil freeze-thaw cycle |
---|---|---|---|---|
土壤冻结初期 Onset of soil freezing period | 0.031 | 0.060 | 0.010 | -0.009 |
土壤冻结期 Soil freezing period | 0.533** | 0.157 | 0.534** | -0.008 |
土壤融化期 Soil thawing period | 0.061 | -0.100 | 0.378* | -0.319 |
生长季节初期 Early growth period | -0.117 | -0.117 | – | -0.431* |
生长季节后期 Later growth period | 0.486** | 0.487** | 0.313 | -0.313 |
冻融季节 Freeze-thaw season | 0.751** | -0.051 | 0.733** | -0.148 |
生长季节 Growth season | 0.151 | 0.133 | 0.027 | 0.029 |
全年 Whole year | 0.589** | 0.582** | 0.547** | -0.023 |
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