植物生态学报 ›› 2008, Vol. 32 ›› Issue (2): 274-286.DOI: 10.3773/j.issn.1005-264x.2008.02.005
叶万辉(), 曹洪麟, 黄忠良, 练琚愉, 王志高, 李林, 魏识广, 王章明
收稿日期:
2007-12-19
接受日期:
2008-01-26
出版日期:
2008-12-19
发布日期:
2008-03-30
通讯作者:
叶万辉
作者简介:
E-mail: why@scbg.ac.cn感谢中国科学院生物多样性委员会对鼎湖山大样地工作的大力支持;感谢加拿大 Alberta大学何芳良教授、台湾东海大学孙义方教授、中国科学院植物研究所马克平教授在建设样地中提出宝贵意见,并在样地选址、调查工作中亲临现场给予指导;感谢美国史密森热带森林研究中心(CFS)生态学家 Richand Condit i教授在数据整理上提供指导和帮助;感谢武汉江韵测绘勘査有限公司在样地建设过程中付出的辛動劳动。同时感谢以下人员在鼎湖山大样地植物调查时给予帮助:华南师范大学的林正媚博士,中国科学院华南植物园的陈炳辉高工、黄玉佳、莫定升、孟泽、张佑昌、吴林芳、叶育石,浙江古田山自然保护区的方腾,中国科学院华南植物园研究生:史军辉、张池、沈浩、李静、穆宏平、韩玉洁;华南师范大学与华南农业大学2002级本科生:施悦谋、陈达丰、谢腾芳、黄婷、陈裕喜、李胜强、陈水莲。
基金资助:
YE Wan-Hui(), CAO Hong-Lin, HUANG Zhong-Liang, LIAN Ju-Yu, WANG Zhi-Gao, LI Lin, WEI Shi-Guang, WANG Zhang-Ming
Received:
2007-12-19
Accepted:
2008-01-26
Online:
2008-12-19
Published:
2008-03-30
Contact:
YE Wan-Hui
摘要:
鼎湖山南亚热带常绿阔叶林是北回归线附近保存较完好的地带性植被,是在南亚热带季风湿润型气候条件下发育的植被类型,具有热带向亚热带的过渡性质,群落结构相对复杂,组成种类相对丰富。参照巴拿马巴洛科罗拉多岛(Barro Colorado Island, BCI) 50 hm2热带雨林样地的技术规范,采用中国森林生物多样性监测网络的统一调查研究方法,于2005年在鼎湖山南亚热带常绿阔叶林建立了20 hm2 固定监测样地,调查并鉴定了样方内胸径(Diameter at breast height, DBH)大于1 cm的木本植物。首次调查结果表明,群落内共有木本植物210种,71 617个活的个体,分属于56科119属。从乔木区系的组成及其特点可以看出,其南亚热带的区系成分占绝对优势,并呈现出由亚热带向热带过渡的趋势。群落垂直结构复杂,地上成层现象较明显,乔木可分为3层,其中重要值最大的锥(Castanopsis chinensis)、木荷(Schima superba)和黄杞(Engelhardtia roxburghiana)均是乔木上层的优势种;中层是群落的主要层,由厚壳桂(Cryptocarya chinensis)、黄叶树(Xanthophyllum hainanense)和华润楠(Machilus chinensis)等中生和耐阴树种组成;下层成分较复杂,物种多样性高,不同地段的物种组成差异较大。样地内物种十分丰富,种-面积曲线拟合显示其物种数量接近于BCI。稀有种比例极高,有110种,占总物种数的52.38%,其中有45%的稀有种源于物种本身的特性,有20%源于区系交汇,人为或自然干扰造成的稀有种占30%以上。样地中所有个体的径级分布(以1 cm等级排列)明显呈倒“J"形,表示群落稳定与正常生长状态。根据对优势种径级结构的分布分析,将各树种的径级结构归纳为4种类型:1)峰型(中径级个体储备型),此类物种为乔木上层优势种;2)倒“J"型 (正常型),此类物种为乔木中层的优势种;3)类倒“J"型(偏正常型),此类物种占据乔木的中、下层;4)“L”型(灌木型),此类物种分布于乔木下层和灌木层。点格局分析得出优势种在各个层上的绝大部分尺度都是聚集分布的,而不同径级的空间分布随物种的不同而呈现出明显的差异性,点格局显示了优势种在相同径级的分布(特别是DBH=10~40 cm)具有一定的空间互补性。对大径级(DBH>40 cm)的504个个体进行空间分布格局分析,结果表明,所有这些个体随机分布于整个样地,但不同的物种在空间分布上存在一定的规律。
叶万辉, 曹洪麟, 黄忠良, 练琚愉, 王志高, 李林, 魏识广, 王章明. 鼎湖山南亚热带常绿阔叶林20公顷样地群落特征研究. 植物生态学报, 2008, 32(2): 274-286. DOI: 10.3773/j.issn.1005-264x.2008.02.005
YE Wan-Hui, CAO Hong-Lin, HUANG Zhong-Liang, LIAN Ju-Yu, WANG Zhi-Gao, LI Lin, WEI Shi-Guang, WANG Zhang-Ming. COMMUNITY STRUCTURE OF A 20 HM2 LOWER SUBTROPICAL EVERGREEN BROADLEAVED FOREST PLOT IN DINGHUSHAN, CHINA. Chinese Journal of Plant Ecology, 2008, 32(2): 274-286. DOI: 10.3773/j.issn.1005-264x.2008.02.005
类群 Group | 组成 Composition | 生长型 Growth type | |||||
---|---|---|---|---|---|---|---|
科Family | 属Genus | 种Species | 乔木Arbor | 小乔木Subarbor | 灌木Shrub | ||
裸子植物Gymnosperm | 2 | 2 | 2 | 2 | - | - | |
被子植物Angiosperm | |||||||
双子叶植物Dicotyledon | 52 | 115 | 206 | 44 | 93 | 69 | |
单子叶植物Monocotyledon | 2 | 2 | 2 | 1 | - | 1 | |
合计 Total | 56 | 119 | 210 | 47 | 93 | 70 |
表1 鼎湖山大样地木本植物统计
Table 1 Statistics of trees in Dinghushan (DHS) plot
类群 Group | 组成 Composition | 生长型 Growth type | |||||
---|---|---|---|---|---|---|---|
科Family | 属Genus | 种Species | 乔木Arbor | 小乔木Subarbor | 灌木Shrub | ||
裸子植物Gymnosperm | 2 | 2 | 2 | 2 | - | - | |
被子植物Angiosperm | |||||||
双子叶植物Dicotyledon | 52 | 115 | 206 | 44 | 93 | 69 | |
单子叶植物Monocotyledon | 2 | 2 | 2 | 1 | - | 1 | |
合计 Total | 56 | 119 | 210 | 47 | 93 | 70 |
科名 Family | 物种数与比率 No. of species (Proportion %) | 科名 Family | 个体数与比率 No. of individuals (Proportion %) |
---|---|---|---|
樟科Lauraceae | 21 (10.00) | 樟科Lauraceae | 14 065 (19.64) |
大戟科Euphorbiaceae | 20 (9.52) | 桃金娘科Myrtaceae | 8 247 (11.52) |
茜草科Rubiaceae | 14 (6.67) | 茜草科Rubiaceae | 8 024 (11.20) |
桑科Moraceae | 11 (5.23) | 野牡丹科Melastomataceae | 5 398 (7.54) |
山茶科Theaceae | 9 (4.29) | 大戟科Euphorbiaceae | 5 114 (7.14) |
桃金娘科Myrtaceae | 9 (4.29) | 杜鹃花科Ericaceae | 4 860 (6.79) |
冬青科Aquifoliaceae | 8 (3.81) | 紫金牛科Myrsinaceae | 4 457 (6.22) |
杜鹃花科Ericaceae | 8 (3.81) | 山茶科Theaceae | 2 881 (4.02) |
杜英科Elaeocarpaceae | 6 (2.86) | 蝶形花科Papilionaceae | 2 791 (3.90) |
蝶形花科Papilionaceae | 5 (2.38) | 壳斗科Fagaceae | 2 604 (3.64) |
总计 Total | 111 (52.86) | 总计 Total | 58 441 (81.60) |
表2 乔木树种物种数和个体数最大的前10个科
Table 2 The top ten families based on the number of species and individuals in Dinghushan (DHS) plot
科名 Family | 物种数与比率 No. of species (Proportion %) | 科名 Family | 个体数与比率 No. of individuals (Proportion %) |
---|---|---|---|
樟科Lauraceae | 21 (10.00) | 樟科Lauraceae | 14 065 (19.64) |
大戟科Euphorbiaceae | 20 (9.52) | 桃金娘科Myrtaceae | 8 247 (11.52) |
茜草科Rubiaceae | 14 (6.67) | 茜草科Rubiaceae | 8 024 (11.20) |
桑科Moraceae | 11 (5.23) | 野牡丹科Melastomataceae | 5 398 (7.54) |
山茶科Theaceae | 9 (4.29) | 大戟科Euphorbiaceae | 5 114 (7.14) |
桃金娘科Myrtaceae | 9 (4.29) | 杜鹃花科Ericaceae | 4 860 (6.79) |
冬青科Aquifoliaceae | 8 (3.81) | 紫金牛科Myrsinaceae | 4 457 (6.22) |
杜鹃花科Ericaceae | 8 (3.81) | 山茶科Theaceae | 2 881 (4.02) |
杜英科Elaeocarpaceae | 6 (2.86) | 蝶形花科Papilionaceae | 2 791 (3.90) |
蝶形花科Papilionaceae | 5 (2.38) | 壳斗科Fagaceae | 2 604 (3.64) |
总计 Total | 111 (52.86) | 总计 Total | 58 441 (81.60) |
分布区类型 Distribution area types | 属数No. of genera | 比率Rate (%) |
---|---|---|
1 世界分布Cosmopolitan | - | - |
2 泛热带分布 Pantropic | 37 | 31.09 |
3 热带亚洲和热带美洲间断分布 Tropical Asia and Tropical America disjuncted | 5 | 4.20 |
4 旧世界热带分布 Old World Tropic | 18 | 15.12 |
5 热带亚洲至热带大洋洲分布 Tropical Asia and Tropical Australasia | 10 | 8.40 |
6 热带亚洲至热带非洲分布 Tropical Asia to Tropical Africa | 8 | 6.72 |
7 热带亚洲分布 Tropical Asia | 29 | 24.37 |
热带成分(分布型2~7) Tropical elements | 107 | 89.92 |
8 北温带分布 North Temperate | 3 | 2.52 |
9 东亚和北美洲间断分布 East Asia and North America disjuncted | 5 | 4.20 |
温带成分(分布型8~9) Temperate elements | 8 | 6.72 |
10 旧世界温带分布 Old World Temperate | - | - |
11 温带亚洲分布 Temperate Asia | - | - |
12 地中海区、西亚至中亚分布 Mediterranean, West Asia to Central Asia | - | - |
13 中亚分布 Central Asia | - | - |
14 东亚分布 East Asia | 4 | 3.36 |
15 中国特有分布 Endemic to China | - | - |
合计 Total | 119 | 100.00 |
表3 鼎湖山大样地木本植物群落种子植物属的分布区类型统计
Table 3 Distribution types of spermatophyte in Dinghushan (DHS) plot
分布区类型 Distribution area types | 属数No. of genera | 比率Rate (%) |
---|---|---|
1 世界分布Cosmopolitan | - | - |
2 泛热带分布 Pantropic | 37 | 31.09 |
3 热带亚洲和热带美洲间断分布 Tropical Asia and Tropical America disjuncted | 5 | 4.20 |
4 旧世界热带分布 Old World Tropic | 18 | 15.12 |
5 热带亚洲至热带大洋洲分布 Tropical Asia and Tropical Australasia | 10 | 8.40 |
6 热带亚洲至热带非洲分布 Tropical Asia to Tropical Africa | 8 | 6.72 |
7 热带亚洲分布 Tropical Asia | 29 | 24.37 |
热带成分(分布型2~7) Tropical elements | 107 | 89.92 |
8 北温带分布 North Temperate | 3 | 2.52 |
9 东亚和北美洲间断分布 East Asia and North America disjuncted | 5 | 4.20 |
温带成分(分布型8~9) Temperate elements | 8 | 6.72 |
10 旧世界温带分布 Old World Temperate | - | - |
11 温带亚洲分布 Temperate Asia | - | - |
12 地中海区、西亚至中亚分布 Mediterranean, West Asia to Central Asia | - | - |
13 中亚分布 Central Asia | - | - |
14 东亚分布 East Asia | 4 | 3.36 |
15 中国特有分布 Endemic to China | - | - |
合计 Total | 119 | 100.00 |
物种名 Species | 多度 Abundance | 分枝数 Ramification | 胸高断面积 Basal area (m2·hm-2) | 重要值 Important value |
---|---|---|---|---|
锥Castanopsis chinensis | 2 311 | 546 | 9.304 | 12.3 |
木荷Schima superba | 2 296 | 260 | 4.132 | 6.6 |
黄杞Engelhardtia roxburghiana | 737 | 149 | 3.475 | 4.8 |
红枝蒲桃Syzygium rehderianum | 5 990 | 347 | 0.879 | 4.7 |
广东金叶子Craibiodendron scleranthum var. kwangtungense | 3 325 | 111 | 1.664 | 4.4 |
香楠Aidia canthioides | 5 996 | 422 | 0.213 | 4.1 |
厚壳桂Cryptocarya chinensis | 2 557 | 1 028 | 1.208 | 3.5 |
黄果厚壳桂Cryptocarya concinna | 4 478 | 106 | 0.177 | 3.3 |
云南银柴Aporusa yunnanensis | 3 747 | 378 | 0.436 | 3.0 |
罗伞树Ardisia quinquegona | 3 702 | 503 | 0.076 | 2.8 |
柏拉木Blastus cochinchinensis | 4 011 | 718 | 0.058 | 2.7 |
肖蒲桃Acmena acuminatissima | 1 484 | 441 | 1.096 | 2.6 |
光叶红豆Ormosia glaberrima | 2 702 | 116 | 0.288 | 2.2 |
黄叶树Xanthophyllum hainanense | 1 873 | 160 | 0.359 | 2.0 |
滇粤山胡椒Lindera metcalfiana | 2 118 | 369 | 0.190 | 1.9 |
肉实树Sarcosperma laurinum | 1 576 | 146 | 0.330 | 1.9 |
华润楠Machilus chinensis | 532 | 65 | 0.845 | 1.7 |
鼎湖钓樟Lindera chunii | 1 302 | 1 016 | 0.151 | 1.5 |
谷木Memecylon ligustrifolium | 1 263 | 74 | 0.092 | 1.4 |
小新木姜Neolitsea umbrosa | 1 352 | 154 | 0.169 | 1.3 |
短序润楠Machilus breviflora | 800 | 327 | 0.344 | 1.3 |
褐叶柄果木Mischocarpus pentapetalus | 1 252 | 95 | 0.044 | 1.3 |
鱼骨木Canthium dicoccum | 603 | 19 | 0.158 | 1.2 |
密花树Rapanea neriifoli | 750 | 56 | 0.289 | 1.2 |
九节Psychotria rubra | 908 | 54 | 0.016 | 1.2 |
毛缘杜鹃Rhododendron henryi var. concavum | 810 | 208 | 0.278 | 1.1 |
马尾松Pinus massonian | 169 | 6 | 0.700 | 1.0 |
岭南山竹子Garcinia oblongifolia | 652 | 48 | 0.104 | 1.0 |
三花冬青Ilex triflora | 642 | 28 | 0.234 | 1.0 |
其它Others | 11 679 | 1 042 | 2.860 | 21.1 |
合计Total | 71 617 | 8 992 | 30.167 | 100.0 |
表4 鼎湖山大样地的物种组成
Table 4 Species composition of live trees ≥ 1.0 cm DBH in Dinghushan (DHS) plot
物种名 Species | 多度 Abundance | 分枝数 Ramification | 胸高断面积 Basal area (m2·hm-2) | 重要值 Important value |
---|---|---|---|---|
锥Castanopsis chinensis | 2 311 | 546 | 9.304 | 12.3 |
木荷Schima superba | 2 296 | 260 | 4.132 | 6.6 |
黄杞Engelhardtia roxburghiana | 737 | 149 | 3.475 | 4.8 |
红枝蒲桃Syzygium rehderianum | 5 990 | 347 | 0.879 | 4.7 |
广东金叶子Craibiodendron scleranthum var. kwangtungense | 3 325 | 111 | 1.664 | 4.4 |
香楠Aidia canthioides | 5 996 | 422 | 0.213 | 4.1 |
厚壳桂Cryptocarya chinensis | 2 557 | 1 028 | 1.208 | 3.5 |
黄果厚壳桂Cryptocarya concinna | 4 478 | 106 | 0.177 | 3.3 |
云南银柴Aporusa yunnanensis | 3 747 | 378 | 0.436 | 3.0 |
罗伞树Ardisia quinquegona | 3 702 | 503 | 0.076 | 2.8 |
柏拉木Blastus cochinchinensis | 4 011 | 718 | 0.058 | 2.7 |
肖蒲桃Acmena acuminatissima | 1 484 | 441 | 1.096 | 2.6 |
光叶红豆Ormosia glaberrima | 2 702 | 116 | 0.288 | 2.2 |
黄叶树Xanthophyllum hainanense | 1 873 | 160 | 0.359 | 2.0 |
滇粤山胡椒Lindera metcalfiana | 2 118 | 369 | 0.190 | 1.9 |
肉实树Sarcosperma laurinum | 1 576 | 146 | 0.330 | 1.9 |
华润楠Machilus chinensis | 532 | 65 | 0.845 | 1.7 |
鼎湖钓樟Lindera chunii | 1 302 | 1 016 | 0.151 | 1.5 |
谷木Memecylon ligustrifolium | 1 263 | 74 | 0.092 | 1.4 |
小新木姜Neolitsea umbrosa | 1 352 | 154 | 0.169 | 1.3 |
短序润楠Machilus breviflora | 800 | 327 | 0.344 | 1.3 |
褐叶柄果木Mischocarpus pentapetalus | 1 252 | 95 | 0.044 | 1.3 |
鱼骨木Canthium dicoccum | 603 | 19 | 0.158 | 1.2 |
密花树Rapanea neriifoli | 750 | 56 | 0.289 | 1.2 |
九节Psychotria rubra | 908 | 54 | 0.016 | 1.2 |
毛缘杜鹃Rhododendron henryi var. concavum | 810 | 208 | 0.278 | 1.1 |
马尾松Pinus massonian | 169 | 6 | 0.700 | 1.0 |
岭南山竹子Garcinia oblongifolia | 652 | 48 | 0.104 | 1.0 |
三花冬青Ilex triflora | 642 | 28 | 0.234 | 1.0 |
其它Others | 11 679 | 1 042 | 2.860 | 21.1 |
合计Total | 71 617 | 8 992 | 30.167 | 100.0 |
种名 Species | 个体数与比率 No. of individuals (Proportion %) |
---|---|
香楠Aidia canthioides | 5 996 (8.37) |
红枝蒲桃Syzygium rehderianum | 5 990 (8.36) |
黄果厚壳桂Cryptocarya concinna | 4 478 (6.25) |
柏拉木Blastus cochinchinensis | 4 011 (5.60) |
云南银柴Aporusa yunnanensis | 3 747 (5.23) |
罗伞树Ardisia quinquegona | 3 702 (5.17) |
广东金叶子Craibiodendron scleranthum var. kwangtungense | 3 325 (4.64) |
光叶红豆Ormosia glaberrima | 2 702 (3.77) |
厚壳桂Cryptocarya chinensis | 2 556 (3.57) |
锥Castanopsis chinensis | 2 312 (3.23) |
合计 Total | 38 819 (54.20) |
表5 个体数最多的10个物种
Table 5 The top ten species based on number of individuals in Dinghushan (DHS) plot
种名 Species | 个体数与比率 No. of individuals (Proportion %) |
---|---|
香楠Aidia canthioides | 5 996 (8.37) |
红枝蒲桃Syzygium rehderianum | 5 990 (8.36) |
黄果厚壳桂Cryptocarya concinna | 4 478 (6.25) |
柏拉木Blastus cochinchinensis | 4 011 (5.60) |
云南银柴Aporusa yunnanensis | 3 747 (5.23) |
罗伞树Ardisia quinquegona | 3 702 (5.17) |
广东金叶子Craibiodendron scleranthum var. kwangtungense | 3 325 (4.64) |
光叶红豆Ormosia glaberrima | 2 702 (3.77) |
厚壳桂Cryptocarya chinensis | 2 556 (3.57) |
锥Castanopsis chinensis | 2 312 (3.23) |
合计 Total | 38 819 (54.20) |
形成原因 Forming factors | 物种数 Species number | 比率 Proportion (%) |
---|---|---|
1. 物种特性 Species characteristics | 49 | 44.54 |
区域稀有种 Rare in Lower Subtropical Area | 32 | 29.09 |
鼎湖山分布较少 Rare in DHS Reserve | 17 | 15.45 |
2. 区系交汇Floristic transitional zone | 24 | 21.82 |
典型亚热带种类 Typical subtropical species | 18 | 16.36 |
沟谷分布种 Ravine species | 6 | 5.46 |
3. 干扰Disturbance | 37 | 31.82 |
阳生性树种 Heliophytes species | 19 | 17.27 |
中生性树种 Mesophyte species | 16 | 14.55 |
果树 Fruit tree | 2 | 1.82 |
合计 Total | 110 | 100.00 |
表6 鼎湖山大样地稀有种形成的原因
Table 6 The forming factors for the rare species in Dinghushan (DHS) plot
形成原因 Forming factors | 物种数 Species number | 比率 Proportion (%) |
---|---|---|
1. 物种特性 Species characteristics | 49 | 44.54 |
区域稀有种 Rare in Lower Subtropical Area | 32 | 29.09 |
鼎湖山分布较少 Rare in DHS Reserve | 17 | 15.45 |
2. 区系交汇Floristic transitional zone | 24 | 21.82 |
典型亚热带种类 Typical subtropical species | 18 | 16.36 |
沟谷分布种 Ravine species | 6 | 5.46 |
3. 干扰Disturbance | 37 | 31.82 |
阳生性树种 Heliophytes species | 19 | 17.27 |
中生性树种 Mesophyte species | 16 | 14.55 |
果树 Fruit tree | 2 | 1.82 |
合计 Total | 110 | 100.00 |
[1] | Bunyavejchewi S, Baker P (1997). Structure, history, and rarity in a seasonal evergreen forest in western Thailand. In: Losos EC, Leigh EG eds. Forest Diversity and Dynamism: Findings from a Network of Large-scale Tropical Forest Plots. University of Chicago Press, Chicago, Illinois, USA, 145-158. |
[2] | Cao HL (曹洪麟), Huang ZL (黄忠良), Zhang LY (张林艳), Kong GH (孔国辉) (2002). Vegetation map of Dinghu Mountain Nature Reserve. Tropical and Subtropical Forest Ecosystem (热带亚热带森林生态系统研究), 9, 1-9. (in Chinese with English abstract) |
[3] | Chase JM (2005). Towards a really unified theory for metacommunities. Functional Ecology, 19, 182-186. |
[4] | Chave J (2004). Neutral theory and community ecology. Ecology Letters, 7, 241-253. |
[5] |
Condit R (1995). Research in large, long-term tropical forest plot. Trends in Ecology & Evolution, 10, 18-23.
URL PMID |
[6] | Condit R (1998). Tropical Forest Census Plots: Methods and Results from Barro Colorado Island, Panama and a Comparison with Other Plots. Springer, Berlin. |
[7] | Condit R, Ashton P, Bunyavejchewin S, Dattaraja HS, Davies S, Esufali S, Ewango C, Foster R, Gunatilleke IA, Gunatilleke CV, Hall P, Harms KE, Hart T, Hernandez C, Hubbell S, Itoh A, Kiratiprayoon S, Lafrankie J, de Lao SL, Makana JR, Noor MN, Kassim AR, Russo S, Sukumar R, Samper C, Suresh HS, Tan S, Thomas S, Valencia R, Vallejo M, Villa G, Zillio T (2006). The importance of demographic niches to tree diversity. Science, 313, 98-101. |
[8] | Condit R, Ashton PS, Baker P, Bunyavejchewin S, Gunatilleke S, Gunatilleke N, Hubbell SP, Foster RB, Itoh A, LaFrankie JV, Lee HS, Losos E, Manokaran N, Sukumar R, Yamakura T (2000). Spatial patterns in the distribution of tropical tree species. Science, 288, 1414-1418. |
[9] |
Condit R, Pitman N, Leigh EG, Chave J, Terborgh J, Foster RB, Núfiez PV, Aguilar S, Valencia R, Villa G, Muller-Landau H, Losos E, Hubbell SP (2002). Beta-diversity in tropical forest trees. Science, 295, 666-669.
URL PMID |
[10] | Dallmeier F, Taylor CM, Mayne JC, Kabel M, Rice R (1992). Case study of SI/MAB biological diversity plot research methodology: effects of hurricane Hugo on the Bisley biodiversity plot, Luquillo Biosphere Preserve, Puerto Rico. In: Dallmeier F ed. Long-term Monitoring of Biological Diversity in Tropical Forest Areas: Methods for Establishment and Inventory of Permanent Plots. MAB Digest 11, UNESCO, Paris, 47-72. |
[11] | He JH (何金海), Chen ZQ (陈兆其), Liang YE (梁永奀) (1982). The soils of Ding Hu Shan biosphere reserve. Tropical and Subtropical Forest Ecosystem (热带亚热带森林生态系统研究), 1, 25-38. (in Chinese with English abstract) |
[12] | Hooper ER, LeGendre P, Condit R (2004). Factors affecting community composition of forest regeneration in deforested, abandoned land in Panama. Ecology, 85, 3313-3326. |
[13] | Huang WF (黄伟峰), Shen XP (沈雪苹) (1982). The climatic ecological environments in the region of Ding Hu Shan. Tropical and Subtropical Forest Ecosystem (热带亚热带森林生态系统研究), 1, 17-24. (in Chinese with English abstract) |
[14] | Huang ZL (黄忠良), Meng ML (蒙满林), Zhang YC (张佑昌) (1998). Climate of Dinghushan Biosphere Reserve. Tropical and Subtropical Forest Ecosystem (热带亚热带森林生态系统研究), 8, 134-139. (in Chinese with English abstract) |
[15] | Hubbell SP (2001). The Unified Neutral Theory of Biodiversity and Biogeography. Princeton University Press, Princeton. |
[16] | Hubbell SP (2006). Neutral theory and the evolution of ecological equivalence. Ecology, 87, 1387-1398. |
[17] | Hubbell SP, Foster RB (1997). Commonness and rarity in a neotropical forest: implications for tropical tree conservation. In: Soule ME ed. Conservation Biology: Science of Scarcity and Diversity. Sinauer Press, Sunderland, Mass, 205-231. |
[18] |
Hubbell SP, Foster RB, O'Brien ST, Harms KE, Condit R, Wechsler B, Wright SJ, de Lao SL (1999). Light gap disturbances, recruitment limitation, and tree diversity in a neotropical forest. Science, 283, 554-557.
URL PMID |
[19] | Kochummen KM, LaFrankie JJV, Manokaran N (1990). Floristic composition of Pasoh forest reserve, a lowland rain forest in peninsular Malaysia. Journal of Tropical Forest Science, 3, 1-3. |
[20] | Kong GH (孔国辉), Ye WH (叶万辉), Huang ZL (黄忠良), Wei P (魏平), Huang YJ (黄玉佳) (1998). Long-term monitoring of the low subtropical evergreen broad-leaved forest in Dinghushan Biosphere Reserve. Ⅰ. Species composition of Castanopsis chinensis, Cryptocarya concinna community and its contribution. Tropical and Subtropical Forest Ecosystem (热带亚热带森林生态系统研究), 8, 1-6. (in Chinese with English abstract) |
[21] | Lee HS, Ashton PS, Ogino K (1995). Long-term ecological research of tropical rain forest in Sarawak. A New Program for Promotion of Basic Sciences Studies of Global Environmental Change with Special Reference to Asia and Pacific Regions 2. Ehime University, Ohzu, Japan. |
[22] | Li L (李林), Zhou XY (周小勇), Huang ZL (黄忠良), Wei SG (魏识广), Shi JH (史军辉)(2006). Study on the relationship between α diversity of plant community and environment on Dinghushan. Acta Ecologica Sinica (生态学报), 26, 2301-2307. (in Chinese with English abstract) |
[23] | Manokaran N, LaFrankie JJV (1990). Stand structure of Pasoh forest reserve, a lowland rain forest in peninsular Malaysia. Journal of Tropical Forest Science, 3, 14-24. |
[24] | McGill BJ (2003). A test of the unified neutral theory of biodiversity. Nature, 422, 881-885. |
[25] | McGill BJ, Maurer BA, Weiser MD (2006). Empirical evaluation of the neutral theory. Ecology, 87, 1411-1423. |
[26] | Peng SJ (彭闪江), Huang ZL (黄忠良), Xu GL (徐国良), Ouyang XJ (欧阳学军), Zhang C (张池)(2003). Effects of habitat heterogeneity on forest community diversity in Dinghushan Biosphere Reserve. Guihaia (广西植物), 23, 391-398. (in Chinese with English abstract) |
[27] | Peng SJ (彭闪江), Xu GL (徐国良)(2005). Seed traits of Castanopsis chinensis and its effects on seed predation patterns in Dinghushan Biosphere Reserve. Ecology and Environment (生态环境), 14, 493-497. (in Chinese with English abstract) |
[28] | Peng SL (彭少麟) (1996). Dynamics of Forest Community in Lower Subtropical Zone (南亚热带森林群落动态学). Science Press, Beijing. (in Chinese) |
[29] | Peng SL (彭少麟), Fang W (方炜) (1994). Studies on the dynamics of dominant populations of Dinghushan forest during successions. Ⅲ. Cryptocarya chinensis and Cryptocarya concinna populations. Journal of Tropical and Subtropical Botany (热带亚热带植物学报), 2, 79-87. (in Chinese with English abstract) |
[30] | Peng SL (彭少麟), Fang W (方炜), Ren H (任海), Huang ZL (黄忠良), Kong GH (孔国辉) (1998). The dynamics on organization in the successional process of Dinghushan Cryptocarya community. Acta Phytoecologica Sinica (植物生态学报), 22, 245-249. (in Chinese with English abstract) |
[31] | Rosenzweig M (1995). Species Diversity in Space and Time. Cambridge University Press, Cambridge, UK. |
[32] | Volkov I, Banavar JR, Hubbell SP, Maritan A (2003). Neutral theory and relative species abundance in ecology. Nature, 424, 1035-1037. |
[33] | Wang ZH (王铸豪), He DQ (何道泉), Song SD (宋绍敦), Chen SP (陈树培), Chen DR (陈定如), Tu MZ (屠梦照) (1982). The vegetation of Ding Hu Shan Biosphere Reserve. Tropical and Subtropical Forest Ecosystem (热带亚热带森林生态系统研究), 1, 77-141. (in Chinese with English abstract) |
[34] | Wu HS (吴厚水), Deng HZ (邓汉增), Chen HT (陈华堂), Zheng LW (郑良文), Liu YD (刘宜钿) (1982). Physico-geographical features of Ding Hu Shan and their dynamic analyses. Tropical and Subtropical Forest Ecosystem (热带亚热带森林生态系统研究), 1, 1-10. (in Chinese with English abstract) |
[35] | Wu ZY (吴征镒) (1991). The areal-types of Chinese genera of seed plants. Acta Botanica Yunnanica (云南植物研究), 4(Suppl.), 1-139. (in Chinese and English) |
[36] | Ye WH (叶万辉) (2000). The maintenance mechanism of plant community and its species diversity. Biodiversity Science (生物多样性), 8, 17-24. (in Chinese with English abstract) |
[37] | Zhang LY (张林艳), Ye WH (叶万辉), Huang ZL (黄忠良) (2006). Assessment of function area design in Dinghushan Biosphere Reserve using landscape ecology principles. Biodiversity Science (生物多样性), 14, 98-106. (in Chinese with English abstract) |
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