植物生态学报 ›› 2007, Vol. 31 ›› Issue (3): 464-469.DOI: 10.17521/cjpe.2007.0057 cstr: 32100.14.cjpe.2007.0057
所属专题: 青藏高原植物生态学:生理生态学; 苔藓生态
收稿日期:2005-12-21
接受日期:2006-06-09
出版日期:2007-12-21
发布日期:2007-05-30
作者简介:* E-mail: wuning@cib.ac.cn基金资助:
WANG Qian, WU Ning*(
), LUO Peng, YI Shao-Liang, BAO Wei-Kai, SHI Fu-Sun
Received:2005-12-21
Accepted:2006-06-09
Online:2007-12-21
Published:2007-05-30
摘要:
利用红油漆标记法,对青藏高原东缘地区壤塘林业局二林场亚高山采伐迹地和云杉(Picea)原始林中的6种藓类近一个生长季的生长速率进行了研究。结合原地同时进行的微气候观测,分析了气候因子和藓类生长速率之间的相关度。这6种藓类中,有5种在原始林和采伐迹地都出现,只有绢藓(Entodon conncinus)仅在采伐迹地出现。不同物种和不同生境条件下藓类的生长速率都不相同。塔藓(Hylocomium splendens)的茎生长速率最大,而阿萨姆曲尾藓(Dicranum assamicum)生长速率最小。生境对塔藓、阿萨姆曲尾藓和细叶羽藓(Thuidium lepidoziaceum)的生长速率影响很大,它们在林内比在采伐迹地生长快。锦丝藓(Actinothuidium hookeri)和垂枝藓(Rhytidiadelphus triquetrus)的生长速率中等并且对生境不敏感。据此可以将藓类分成生境敏感型和不敏感型。微气候5~7月分析显示林内比采伐迹地的辐射通量低,并且更为干燥。但是早上林内的蒸汽压亏缺(Vapor pressure deficit, VPD)一直比采伐迹地低。林内较低的蒸汽压亏缺和较低的辐射通量使其成为更有利于藓类的生长场所。对于那些生境类型不敏感的藓类,微地形的效应也许抵消了这种大生境的效应。藓类的生理生态特征如变水(Poikilohydry)特征等对其在严酷气候条件下的生存和生长起着重要的作用。如同积温一样,藓类的生长速率同样可以指示生境的适宜度, 因为其生长和蒸汽压亏缺紧密相关,是温度和湿度的函数,而这两个因素对于川西亚高山地区森林人工更新时幼苗的建植极为关键。因此藓类的生长状况可以作为指示适宜植树生境的指标。
王乾, 吴宁, 罗鹏, 易绍良, 包维楷, 石福孙. 青藏高原东缘亚高山针叶林和采伐迹地中藓类生长速率及其影响因子. 植物生态学报, 2007, 31(3): 464-469. DOI: 10.17521/cjpe.2007.0057
WANG Qian, WU Ning, LUO Peng, YI Shao-Liang, BAO Wei-Kai, SHI Fu-Sun. MOSS GROWTH RATE AND ITS ENVIRONMENTAL DETERMINANTS IN SUBALPINE CONIFEROUS FOREST AND CLEAR-CUT LAND IN EASTERN TIBETAN PLATEAU, CHINA. Chinese Journal of Plant Ecology, 2007, 31(3): 464-469. DOI: 10.17521/cjpe.2007.0057
| 变量 Variable | 5月平均值 Mean in May | 7月平均值 Mean in July | 5月标准方差 SD in May | 7月标准方差 SD in July | 5月May | 7月July |
|---|---|---|---|---|---|---|
| p | ||||||
| 采伐迹地气温CAT (℃) | 6.18 | 9.05 | 4.54 | 5.09 | ||
| 林地气温FAT (℃) | 2.15 | 8.49 | 3.74 | 4.32 | <0.001 | 0.009 |
| 采伐迹地相对湿度CRH (%) | 71.44 | 74.49 | 23.9 | 23.5 | ||
| 林内相对湿度FRH (%) | 69.51 | 59.36 | 24.5 | 23.8 | 0.026 | <0.001 |
| 采伐迹地VPD VPDC (KPa) | 0.344 | 0.371 | 0.369 | 0.389 | ||
| 林内VPD VPDF (KPa) | 0.276 | 0.513 | 0.302 | 0.417 | <0.001 | <0.001 |
| 采伐迹地VPD VPDC (6:00~11:00) (KPa) | 0.431 | 0.417 | 0.266 | 0.295 | ||
| 林内VPD VPDF (6:00~11:00) (KPa) | 0.241 | 0.331 | 0.232 | 0.228 | <0.001 | <0.001 |
表1 5和7月气象资料单因素方差分析
Table 1 One-way ANOVA analysis of May and July data
| 变量 Variable | 5月平均值 Mean in May | 7月平均值 Mean in July | 5月标准方差 SD in May | 7月标准方差 SD in July | 5月May | 7月July |
|---|---|---|---|---|---|---|
| p | ||||||
| 采伐迹地气温CAT (℃) | 6.18 | 9.05 | 4.54 | 5.09 | ||
| 林地气温FAT (℃) | 2.15 | 8.49 | 3.74 | 4.32 | <0.001 | 0.009 |
| 采伐迹地相对湿度CRH (%) | 71.44 | 74.49 | 23.9 | 23.5 | ||
| 林内相对湿度FRH (%) | 69.51 | 59.36 | 24.5 | 23.8 | 0.026 | <0.001 |
| 采伐迹地VPD VPDC (KPa) | 0.344 | 0.371 | 0.369 | 0.389 | ||
| 林内VPD VPDF (KPa) | 0.276 | 0.513 | 0.302 | 0.417 | <0.001 | <0.001 |
| 采伐迹地VPD VPDC (6:00~11:00) (KPa) | 0.431 | 0.417 | 0.266 | 0.295 | ||
| 林内VPD VPDF (6:00~11:00) (KPa) | 0.241 | 0.331 | 0.232 | 0.228 | <0.001 | <0.001 |
图1 计算的5月(A)和7月(B)林内与采伐迹地连续多日蒸气压亏缺(VPD)差异日动态
Fig.1 Calculated daily course of vapor pressure deficit (VPD) difference from consecutive days between forest interior and clear-cut in May (A) and July (B)
图2 采伐迹地和林内的藓类在生长季期间平均生长速率(平均生长速率+标准偏差)
Fig.2 The bar graph of mosses mean growth rate at clear-cut land and forest interior in growing season (mean growth rate + SD) 1. 锦丝藓 Actinothuidium hookeri 2. 垂枝藓 Rhytidiadelphus triquetrus 3. 绢藓 Entodon conncinus 4. 塔藓 Hylocomium splendens 5. 阿萨姆曲尾藓 Dicranum assamicum 6. 细叶羽藓 Thuidium lepidoziaceum
| 物种 Species | 生境 Habitat | 平均值 Mean | 均方差 SD | 数量 Number | 偏度 Skewness | 峰度 Kurtosis | F | p |
|---|---|---|---|---|---|---|---|---|
| 锦丝藓Actinothuidium hookeri | C | 0.058 | 0.023 | 23 | 0.103 | 2.20 | 3.90 | 0.052 |
| F | 0.070 | 0.024 | 50 | 0.802 | 4.47 | |||
| 垂枝藓Rhytidiadelphus triquetrus | C | 0.032 | 0.025 | 94 | 0.535 | 2.61 | 1.19 | 0.28 |
| F | 0.035 | 0.024 | 215 | 0.961 | 6.20 | |||
| 绢藓Entodon conncinus | C | 0.029 | 0.033 | 166 | 1.646 | 6.33 | - | - |
| F | - | - | - | - | - | |||
| 塔藓Hylocomium splendens | C | 0.020 | 0.020 | 25 | 0.048 | 1.42 | 21.02 | <0.001 |
| F | 0.142 | 0.059 | 215 | 0.398 | 3.78 | |||
| 阿萨姆曲尾藓Dicranum assamicum | C | 0.004 | 0.014 | 141 | 3.366 | 13.57 | 49.52 | <0.001 |
| F | 0.023 | 0.021 | 55 | 0.125 | 1.64 | |||
| 细叶羽藓Thuidium lepidoziaceum | C | 0.053 | 0.032 | 82 | 0.367 | 3.20 | 14.94 | <0.001 |
| F | 0.072 | 0.035 | 100 | 1.109 | 4.79 |
表2 藓类在不同生境中的平均生长速率单因素方差分析
Table 2 One-way ANOVA of the mean growth rate of mosses in habitats
| 物种 Species | 生境 Habitat | 平均值 Mean | 均方差 SD | 数量 Number | 偏度 Skewness | 峰度 Kurtosis | F | p |
|---|---|---|---|---|---|---|---|---|
| 锦丝藓Actinothuidium hookeri | C | 0.058 | 0.023 | 23 | 0.103 | 2.20 | 3.90 | 0.052 |
| F | 0.070 | 0.024 | 50 | 0.802 | 4.47 | |||
| 垂枝藓Rhytidiadelphus triquetrus | C | 0.032 | 0.025 | 94 | 0.535 | 2.61 | 1.19 | 0.28 |
| F | 0.035 | 0.024 | 215 | 0.961 | 6.20 | |||
| 绢藓Entodon conncinus | C | 0.029 | 0.033 | 166 | 1.646 | 6.33 | - | - |
| F | - | - | - | - | - | |||
| 塔藓Hylocomium splendens | C | 0.020 | 0.020 | 25 | 0.048 | 1.42 | 21.02 | <0.001 |
| F | 0.142 | 0.059 | 215 | 0.398 | 3.78 | |||
| 阿萨姆曲尾藓Dicranum assamicum | C | 0.004 | 0.014 | 141 | 3.366 | 13.57 | 49.52 | <0.001 |
| F | 0.023 | 0.021 | 55 | 0.125 | 1.64 | |||
| 细叶羽藓Thuidium lepidoziaceum | C | 0.053 | 0.032 | 82 | 0.367 | 3.20 | 14.94 | <0.001 |
| F | 0.072 | 0.035 | 100 | 1.109 | 4.79 |
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