不同龄级划分方法对种群存活分析的影响——以水灾迹地油松和华山松种群生存分析为例

doi:10.3773/j.issn.1005-264x.2008.02.024

摘要/Abstract

摘要：

生命表、存活曲线和生存分析是研究种群动态的重要工具,其关键就是科学正确地划分龄级。该文研究了水灾迹地不同龄级划分方法对油松(Pinus tabulaeformis)和华山松(P. armandii)种群存活曲线和生存分析函数曲线的影响。华山松以实际年龄作为龄级划分指标时,存活曲线为Ⅱ型;地径和树高作为划分指标时,存活曲线为Ⅲ型。生存函数和累计死亡率函数也发生类似的类型变化。油松实际年龄、地径和树高作为龄级的划分指标时,存活曲线(Ⅱ型)、生存率函数和累计死亡率函数类型一致,均为直线型。死亡率函数和危险率函数在不同树种和不同龄级划分方法间不存在明显差异。因此,华山松地径和树高作为龄级划分的代用指标不可行,而油松可行。其主要原因是华山松树高和地径与年龄的关系为指数函数关系,而油松则为直线函数关系。在此基础上把华山松地径和树高取自然对数后重新划分龄级,则存活曲线关系在地径、树高和实际年龄之间不存在差异,均为Ⅱ型。从而得出结论,地径、树高和年龄之间线性关系与否是地径和树高作为实际年龄代用指标可行性和准确性的关键。因此,在植物种群统计研究中,大小作为龄级划分代用指标要慎用。在未来研究中,种群个体年龄和大小关系及其影响因素的研究对植物种群数量动态分析和种群统计学的发展具有重要意义。

关键词: 秦岭北坡, 种群, 静态生命表, 油松, 华山松

Abstract:

Aims Our objectives were to 1) explore the feasibility of using individual height and basal diameter as substitutes for age, and 2) determine the influence of using these parameters on survival curve analysis for Pinus armandii and P. tabulaeformis populations in an area damaged by water disaster in the northern Qinling Mountain.
Methods We investigated two dominant species, P. tabulaeformis and P. armandii, which had regenerated after damage by water disaster 17 years earlier. We measured height, basal diameter and age of two species in 6 plots (total plot area was 11 900 m ²). We determined age by counting number of whorls of branches.
Important findings It was infeasible to use basal diameter and tree height in place of age for P. armandii because the relationship to age was an exponential function for both parameters; therefore, we used logarithmic values for basal diameter and tree height. In contrast, the functions were linear for P. tabulaeformis. The survival curves were all linear type with no type differences between base diameter, tree height and factual age. Our study indicates that it is feasible to use basal diameter and tree height as indices for factual age when the relationships of basal diameter and tree height to age are linear. Investigation of relations of age and size and their influential factors is important for the analysis of plant population demography.

Key words: Qinling Mountain, population, static life table, Pinus armandii, P. tabulaeformis

何亚平, 费世民, 蒋俊明, 陈秀明, 张旭东, 何飞. 不同龄级划分方法对种群存活分析的影响——以水灾迹地油松和华山松种群生存分析为例. 植物生态学报, 2008, 32(2): 448-455. DOI: 10.3773/j.issn.1005-264x.2008.02.024

HE Ya-Ping, FEI Shi-Min, JIANG Jun-Ming, CHEN Xiu-Ming, ZHANG Xu-Dong, HE Fei. INFLUENCES OF SUBSTITUTING SIZE VARIABLES FOR AGE ON POPULATION SURVIVAL ANALYSIS: A CASE STUDY OF PINUS TABULAEFORMIS AND P. ARMANDII IN QINLING MOUNTAIN, CHINA. Chinese Journal of Plant Ecology, 2008, 32(2): 448-455. DOI: 10.3773/j.issn.1005-264x.2008.02.024

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链接本文: https://www.plant-ecology.com/CN/10.3773/j.issn.1005-264x.2008.02.024

https://www.plant-ecology.com/CN/Y2008/V32/I2/448

图/表 6

图1 水灾迹地油松和华山松种群的存活曲线

Fig.1 Survival curves of Pinus tabulaeformis and P. armandii populations regenerated following the water disaster

图2 水灾迹地油松和华山松种群的生存率函数

Fig.2 S(i) of Pinus tabulaeformis and P. armandii populations regerterated following the water disaster

图3 水灾迹地油松和华山松种群的累计死亡率函数

Fig.3 F(i) of Pinus tabulaeformis and P. armandii population regerterated following the water disaster

图4 水灾迹地油松和华山松种群的死亡密度函数和危险率函数

Fig.4 F(ti) and λ(i) of Pinus tabulaeformis and P. armandii population regerterated following the water disaster

图5 水灾迹地油松和华山松树高、地径和年龄的关系

Fig.5 The relation of height and base diameter of Pinus tabulaeformis and P. armandii to age in the area damaged by the water disaster

图6 水灾迹地华山松种群存活曲线(经校正)

Fig.6 Survival curves of Pinus armandii populations regenerated following the water disaster (corrected)

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