小兴安岭7种典型林型林分生物量碳密度与固碳能力

doi:10.17521/cjpe.2015.0014

植物生态学报 ›› 2015, Vol. 39 ›› Issue (2): 140-158.DOI: 10.17521/cjpe.2015.0014

小兴安岭7种典型林型林分生物量碳密度与固碳能力

胡海清¹, 罗碧珍¹, 魏书精^1,^2,³, 魏书威⁴, 孙龙¹, 罗斯生¹, 马洪斌^1,^*()

¹东北林业大学林学院, 哈尔滨 150040
²桂林理工大学土木与建筑工程学院, 广西桂林 541004
³广西建筑新能源与节能重点实验室, 广西桂林 541004
⁴兰州理工大学设计艺术学院, 兰州 730050

收稿日期:2014-08-11 接受日期:2014-12-01 出版日期:2015-02-01 发布日期:2015-03-10
通讯作者: 马洪斌
作者简介:
# 共同第一作者
基金资助:
黑龙江省博士后基金(LBH-Z13012)、广西自然科学基金(2014GXNSFBA118108)、国家公益性行业科研重大专项(201404402)、国家科技支撑计划项目(2011BAD37B01)、国家自然科学基金(51208244和31070544)、黑龙江省科技攻关重点项目(GA09B201-06)和国家重点基础研究发展计划(2011CB403203)、林业公益性行业科研专项(2010- 04003-6)

Biomass carbon density and carbon sequestration capacity in seven typical forest types of the Xiaoxing’an Mountains, China

HU Hai-Qing¹, LUO Bi-Zhen¹, WEI Shu-Jing^1,^2,³, WEI Shu-Wei⁴, SUN Long¹, LUO Si-Sheng¹, MA Hong-Bin^1,^*()

¹College of Forestry, Northeast Forestry University, Harbin 150040, China
²College of Civil Engineering and Architecture, Guilin University of Technology, Guilin, Guangxi 541004, China
³Guangxi Key Laboratory of New Energy and Building Energy Saving, Guilin, Guangxi 541004, China
⁴College of Design Art, Lanzhou University of Technology, Lanzhou 730050, China

Received:2014-08-11 Accepted:2014-12-01 Online:2015-02-01 Published:2015-03-10
Contact: Hong-Bin MA
About author:
# Co-first authors

摘要/Abstract

摘要：

森林生物碳储量作为森林生态系统碳库的重要组成部分, 在全球碳循环中发挥着重要作用。以小兴安岭7种典型林型为研究对象, 通过外业样地调查与室内实验分析相结合的方法, 从林分尺度对林分生物量与碳密度进行计量, 分析了林分生物碳储量的空间分配格局, 并对林分年固碳能力与碳汇潜力进行了探讨。结果表明: 小兴安岭不同林型从幼龄林到成熟林的乔木层碳密度增长速率为: 蒙古栎(Quercus mongolica)林>兴安落叶松(Larix gmelinii)林>云冷杉(Picea-Abies)林>樟子松(Pinus sylvestris var. mongolica)林>山杨(Populus davidiana)林>红松(Pinus koraiensis)林>白桦(Betula platyphylla)林。7种典型林型不同龄组(幼龄林、中龄林、近熟林和成熟林)林分生物量碳密度分别为: 红松林31.4、74.7、118.4和130.2 t·hm^-2; 兴安落叶松林28.9、44.3、74.2和113.3 t·hm^-2; 樟子松林22.8、52.0、71.1和92.6 t·hm^-2; 云冷杉林23.1、44.1、77.6和130.3 t·hm^-2; 白桦林18.8、35.3、66.6和88.5 t·hm^-2; 蒙古栎林25.0、20.0、47.5和68.9 t·hm^-2; 山杨林19.8、28.7、43.7和76.6 t·hm^-2。红松林、兴安落叶松林、樟子松林和蒙古栎林在幼龄林时林分年固碳量较高, 其他林型在成熟林时林分年固碳量较高。7种典型林型不同龄组的林分生物量碳密度均随林龄增长而增加, 但不同林型的碳汇功能存在差异, 同一林型不同林龄的生物量碳密度增幅差异也较大。林分年固碳量在0.4-2.8 t·hm^-2之间, 碳汇能力较强、碳汇潜力较大。尤其是小兴安岭目前林分质量较差, 幼龄林和中龄林所占的比重较大, 具有较大的碳汇潜力。研究结果可为森林经营管理及碳汇功能评价提供参考。

关键词: 林分生物量, 碳密度, 固碳能力, 年固碳量, 碳汇管理路径, 小兴安岭

Abstract: Aims

Forests are the world’s largest carbon (C) pool and sink among the terrestrial ecosystems. The amount of C in vegetation plays an important role in the global C cycle and balance. Our objectives were to assess C density and sequestration capacity in seven typical forest types of the Xiaoxing’an Mountains, Heilongjiang Province, China and to understand the implication of the C sink to the regional C budget and future forest C management.

Methods

Field surveys were combined with laboratory analysis and allometric equations for obtaining data for a variety of variables. Seven typical forest types in the Xiaoxing’an Mountains were studied based on age groups and plant functional groups (trees, shrubs, herbaceous and litter), including Pinus koraiensis, Larix gmelinii, Pinus sylvestris var. mongolica, Picea-Abies, Betula platyphylla, Quercus mongolica, and Populus davidiana forests. Surveys were made on C density and annual carbon gains in trees, understory shrubs, herbaceous plants and litter for each forest type. The forest stands were classified into age groups for estimating the biomass and C density of the study area.

Important findings

The C density of the seven forest types in different age groups varied widely. The C density per unit area for young, middle-aged, near mature and mature forests of each forest type were as follows: 31.4, 74.7, 118.4 and 130.2 t·hm^-2 in Pinus koraiensis; 28.9, 44.3, 74.2 and 113.3 t·hm^-2 in L. gmelinii; 22.8, 52.0, 71.1 and 92.6 t·hm^-2 in Pinus sylvestris var. mongolica; 23.1, 44.1, 77.6 and 130.3 t·hm^-2 in Picea-Abies; 18.8, 35.3, 66.6 and 88.5 t·hm^-2 in B. platyphylla; 25.0, 20.0, 47.4 and 68.9 t·hm^-2 in Q. mongolica; and 19.8, 28.7, 43.7 and 76.6 t·hm^-2 in Populus davidiana forests, respectively. These results show that biomass C stocks in the Xiao- xing’an Mountains play an important role in the C cycle and regional C balance. Different forest types and stands of different age groups varied greatly in C stocks. Because most growth in the seven forest types occurs in the young and middle-aged forest stands, these age groups are considered to have a great potential to increase the biomass C density. This significant C sink will be further enhanced in the Xiaoxing’an Mountains with the development and restoration designed to provide specific ecological services including C sequestration.

Key words: forest stand biomass, carbon density, carbon sequestration capacity, annual carbon sequestration, carbon sink management path, Xiaoxing’an Mountains

胡海清, 罗碧珍, 魏书精, 魏书威, 孙龙, 罗斯生, 马洪斌. 小兴安岭7种典型林型林分生物量碳密度与固碳能力. 植物生态学报, 2015, 39(2): 140-158. DOI: 10.17521/cjpe.2015.0014

HU Hai-Qing,LUO Bi-Zhen,WEI Shu-Jing,WEI Shu-Wei,SUN Long,LUO Si-Sheng,MA Hong-Bin. Biomass carbon density and carbon sequestration capacity in seven typical forest types of the Xiaoxing’an Mountains, China. Chinese Journal of Plant Ecology, 2015, 39(2): 140-158. DOI: 10.17521/cjpe.2015.0014

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图/表 7

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林型 Forest type	龄组划分 Age group
林型 Forest type	幼龄林 Young forest (a)	中龄林 Middle forest (a)	近熟林 Premature forest (a)	成熟林 Mature forest (a)
红松林 Pinus koraiensis forest	≤60	61-100	101-120	120-160
兴安落叶松林 Larix gmelinii forest	≤40	41-80	81-100	101-140
樟子松林 Pinus sylvestris var. mongolica forest	≤40	41-80	81-100	101-140
云冷杉林 Picea-Abies forest	≤60	61-100	101-120	120-160
白桦林 Betula platyphylla forest	≤30	31-50	51-60	61-80
蒙古栎林 Quercus mongolica forest	≤40	41-60	61-80	81-120
山杨林 Populus davidiana forest	≤10	11-15	16-20	21-30

林型 Forest type	龄组划分 Age group
林型 Forest type	幼龄林 Young forest (a)	中龄林 Middle forest (a)	近熟林 Premature forest (a)	成熟林 Mature forest (a)
红松林 Pinus koraiensis forest	≤60	61-100	101-120	120-160
兴安落叶松林 Larix gmelinii forest	≤40	41-80	81-100	101-140
樟子松林 Pinus sylvestris var. mongolica forest	≤40	41-80	81-100	101-140
云冷杉林 Picea-Abies forest	≤60	61-100	101-120	120-160
白桦林 Betula platyphylla forest	≤30	31-50	51-60	61-80
蒙古栎林 Quercus mongolica forest	≤40	41-60	61-80	81-120
山杨林 Populus davidiana forest	≤10	11-15	16-20	21-30

林型 Forest type	组分 Component	a	b	决定系数 Coefficient of determination (R²)	样本数 No. of samples (N)
红松林 Pinus koraiensis forest	干 Trunk	0.046 65	0.902 37	0.97	30
	枝 Branch	0.031 23	0.612 48	0.99	30
	叶 Leaf	0.012 35	0.643 47	0.95	30
	皮 Bark	0.029 57	0.564 89	0.90	30
	根 Root	0.009 25	0.739 65	0.96	30
兴安落叶松林 Larix gmelinii forest	干 Trunk	0.013 02	1.021 54	0.99	30
	枝 Branch	0.001 29	1.039 98	0.92	30
	叶 Leaf	0.011 12	0.637 45	0.95	30
	皮 Bark	0.031 27	0.615 24	0.97	30
	根 Root	0.041 25	0.684 51	0.99	30
樟子松林 Pinus sylvestris var. mongolica forest	干 Trunk	0.051 41	0.862 14	0.94	30
	枝 Branch	0.014 63	0.724 58	0.97	30
	叶 Leaf	0.021 24	0.524 78	0.91	30
	皮 Bark	0.057 81	0.442 87	0.92	30
	根 Root	0.026 31	0.686 56	0.95	30
云冷杉林 Picea-Abies forest	干 Trunk	0.322 91	0.679 19	0.97	30
	枝 Branch	0.022 57	0.635 64	0.90	30
	叶 Leaf	0.004 18	0.821 35	0.92	30
	皮 Bark	0.024 21	0.572 15	0.94	30
	根 Root	0.011 54	0.812 34	0.94	30
白桦林 Betula platyphylla forest	干 Trunk	0.011 75	1.102 52	0.97	30
	枝 Branch	0.010 24	0.805 47	0.97	30
	叶 Leaf	0.013 47	0.649 47	0.90	30
	皮 Bark	0.024 69	0.631 01	0.92	30
	根 Root	0.048 87	0.632 46	0.96	30
蒙古栎林 Quercus mongolica forest	干 Trunk	0.011 97	1.092 48	0.91	30
	枝 Branch	0.008 45	0.894 18	0.90	30
	叶 Leaf	0.006 24	0.828 54	0.91	30
	皮 Bark	0.008 72	0.817 59	0.92	30
	根 Root	0.010 54	0.835 38	0.93	30
山杨林 Populus davidiana forest	干 Trunk	0.235 14	0.853 24	0.93	30
	枝 Branch	0.021 54	0.862 15	0.98	30
	叶 Leaf	0.009 79	0.856 14	0.96	30
	皮 Bark	0.052 23	0.632 17	0.97	30
	根 Root	0.125 87	0.558 74	0.98	30

林型 Forest type	组分 Component	a	b	决定系数 Coefficient of determination (R²)	样本数 No. of samples (N)
红松林 Pinus koraiensis forest	干 Trunk	0.046 65	0.902 37	0.97	30
	枝 Branch	0.031 23	0.612 48	0.99	30
	叶 Leaf	0.012 35	0.643 47	0.95	30
	皮 Bark	0.029 57	0.564 89	0.90	30
	根 Root	0.009 25	0.739 65	0.96	30
兴安落叶松林 Larix gmelinii forest	干 Trunk	0.013 02	1.021 54	0.99	30
	枝 Branch	0.001 29	1.039 98	0.92	30
	叶 Leaf	0.011 12	0.637 45	0.95	30
	皮 Bark	0.031 27	0.615 24	0.97	30
	根 Root	0.041 25	0.684 51	0.99	30
樟子松林 Pinus sylvestris var. mongolica forest	干 Trunk	0.051 41	0.862 14	0.94	30
	枝 Branch	0.014 63	0.724 58	0.97	30
	叶 Leaf	0.021 24	0.524 78	0.91	30
	皮 Bark	0.057 81	0.442 87	0.92	30
	根 Root	0.026 31	0.686 56	0.95	30
云冷杉林 Picea-Abies forest	干 Trunk	0.322 91	0.679 19	0.97	30
	枝 Branch	0.022 57	0.635 64	0.90	30
	叶 Leaf	0.004 18	0.821 35	0.92	30
	皮 Bark	0.024 21	0.572 15	0.94	30
	根 Root	0.011 54	0.812 34	0.94	30
白桦林 Betula platyphylla forest	干 Trunk	0.011 75	1.102 52	0.97	30
	枝 Branch	0.010 24	0.805 47	0.97	30
	叶 Leaf	0.013 47	0.649 47	0.90	30
	皮 Bark	0.024 69	0.631 01	0.92	30
	根 Root	0.048 87	0.632 46	0.96	30
蒙古栎林 Quercus mongolica forest	干 Trunk	0.011 97	1.092 48	0.91	30
	枝 Branch	0.008 45	0.894 18	0.90	30
	叶 Leaf	0.006 24	0.828 54	0.91	30
	皮 Bark	0.008 72	0.817 59	0.92	30
	根 Root	0.010 54	0.835 38	0.93	30
山杨林 Populus davidiana forest	干 Trunk	0.235 14	0.853 24	0.93	30
	枝 Branch	0.021 54	0.862 15	0.98	30
	叶 Leaf	0.009 79	0.856 14	0.96	30
	皮 Bark	0.052 23	0.632 17	0.97	30
	根 Root	0.125 87	0.558 74	0.98	30

林型 Forest type	龄组 Age group	平均林龄 Average forest age (a)	平均胸径 Average diameter at breast height (cm)	平均树高 Average tree height (m)
红松林 Pinus koraiensis forest	幼龄林 Young forest	24.23 ± 2.21	11.94 ± 1.34	9.95 ± 0.97
	中龄林 Middle-aged forest	68.54 ± 7.24	17.66 ± 2.45	16.54 ± 3.74
	近熟林 Near mature forest	109.19 ± 3.26	20.48 ± 2.18	21.62 ± 5.86
	成熟林 Mature forest	123.50 ± 12.34	21.31 ± 5.67	22.46 ± 4.59
兴安落叶松林 Larix gmelinii forest	幼龄林 Young forest	21.66 ± 1.78	9.46 ± 1.15	9.44 ± 0.91
	中龄林 Middle-aged forest	63.78 ± 10.51	13.51 ± 2.30	12.13 ± 3.48
	近熟林 Near mature forest	84.50 ± 2.96	17.12 ± 6.47	17.59 ± 3.17
	成熟林 Mature forest	135.47 ± 7.28	21.04 ± 6.19	20.27 ± 5.89
樟子松林 Pinus sylvestris var. mongolica forest	幼龄林 Young forest	25.48 ± 3.12	9.11 ± 1.62	8.83 ± 1.45
	中龄林 Middle-aged forest	62.53 ± 8.84	16.77 ± 3.57	15.38 ± 2.84
	近熟林 Near mature forest	89.18 ± 4.16	18.91 ± 3.66	17.01 ± 7.11
	成熟林 Mature forest	122.45 ± 10.47	21.31 ± 6.41	19.44 ± 6.97
云冷杉林 Picea-Abies forest	幼龄林 Young forest	29.60 ± 4.15	7.26 ± 1.01	9.77 ± 1.22
	中龄林 Middle-aged forest	68.00 ± 6.74	11.89 ± 1.34	13.03 ± 1.42
	近熟林 Near mature forest	111.97 ± 3.48	16.57 ± 3.59	16.02 ± 3.47
	成熟林 Mature forest	135.46 ± 15.42	21.54 ± 2.49	22.75 ± 5.64
白桦林 Betula platyphylla forest	幼龄林 Young forest	19.73 ± 3.15	9.55 ± 1.32	10.31 ± 1.46
	中龄林 Middle-aged forest	42.93 ± 5.85	11.63 ± 2.64	13.59 ± 3.69
	近熟林 Near mature forest	56.27 ± 1.42	14.99 ± 4.59	15.03 ± 3.67
	成熟林 Mature forest	74.81 ± 3.00	16.29 ± 5.67	18.04 ± 6.68
蒙古栎林 Quercus mongolica forest	幼龄林 Young forest	23.87 ± 4.25	7.53 ± 0.98	5.87 ± 0.89
	中龄林 Middle-aged forest	48.00 ± 3.33	11.30 ± 1.28	7.15 ± 0.78
	近熟林 Near mature forest	66.59 ± 4.49	14.72 ± 1.94	12.38 ± 2.00
	成熟林 Mature forest	101.42 ± 10.29	17.03 ± 2.37	13.43 ± 2.79
山杨林 Populus davidiana forest	幼龄林 Young forest	8.64 ± 0.54	5.47 ± 0.24	5.56 ± 0.51
	中龄林 Middle-aged forest	13.75 ± 0.48	6.79 ± 0.84	7.84 ± 0.36
	近熟林 Near mature forest	19.53 ± 0.89	8.73 ± 0.74	9.25 ± 0.49
	成熟林 Mature forest	27.61 ± 1.12	11.39 ± 1.37	10.76 ± 0.87

小兴安岭7种典型林型林分生物量碳密度与固碳能力

Biomass carbon density and carbon sequestration capacity in seven typical forest types of the Xiaoxing’an Mountains, China

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林型 Forest type	龄组 Age group	乔木层 Tree						灌木层 Shrub layer	草本层 Herbaceous layer	凋落物层 Litter layer	林分生物量 Stand biomass
林型 Forest type	龄组 Age group	干 Trunk	枝 Branch	叶 Leaf	皮 Bark	根 Root	小计 Total	灌木层 Shrub layer	草本层 Herbaceous layer	凋落物层 Litter layer	林分生物量 Stand biomass
红松林 Pinus koraiensis forest	幼龄林 Young forest	32.58 ± 3.27	2.66 ± 0.24	1.32 ± 0.21	1.78 ± 0.45	1.98 ± 0.42	40.33 ± 3.59	10.57 ± 2.16	3.24 ± 0.68	8.99 ± 2.03	63.13 ± 4.26
	中龄林 Middle-aged forest	104.45 ± 11.49	5.87 ± 1.03	3.02 ± 0.57	3.70 ± 0.89	5.15 ± 0.67	122.19 ± 13.45	12.34 ± 1.85	4.57 ± 0.94	9.54 ± 1.18	148.64 ± 15.68
	近熟林 Near mature forest	173.78 ± 22.37	8.29 ± 1.55	4.35 ± 1.34	5.09 ± 1.32	7.82 ± 1.23	199.32 ± 26.34	15.24 ± 1.97	5.21 ± 0.86	10.55 ± 1.69	230.32 ± 28.56
	成熟林 Mature forest	193.23 ± 19.57	8.91 ± 1.33	4.69 ± 0.76	5.44 ± 1.42	8.53 ± 1.67	220.79 ± 21.45	8.24 ± 1.59	5.57 ± 0.74	18.57 ± 2.87	253.17 ± 22.43
兴安落叶松林 Larix gmelinii forest	幼龄林 Young forest	12.72 ± 1.69	1.43 ± 0.17	0.82 ± 0.37	1.98 ± 0.34	4.16 ± 0.86	21.09 ± 2.26	18.66 ± 4.19	7.59 ± 1.47	10.21 ± 2.46	57.55 ± 2.85
	中龄林 Middle-aged forest	34.03 ± 4.58	3.89 ± 0.78	1.51 ± 0.56	3.57 ± 0.86	8.04 ± 1.27	51.04 ± 5.26	9.58 ± 1.51	11.28 ± 1.39	14.59 ± 3.45	86.49 ± 6.97
	近熟林 Near mature forest	80.70 ± 16.74	9.36 ± 1.67	2.59 ± 0.34	6.01 ± 1.26	14.34 ± 2.19	112.99 ± 18.75	4.24 ± 0.24	8.84 ± 1.75	15.37 ± 3.31	141.44 ± 20.17
	成熟林 Mature forest	142.14 ± 25.67	16.66 ± 3.49	3.68 ± 0.81	8.46 ± 1.78	20.95 ± 2.87	191.88 ± 25.79	2.27 ± 0.15	3.21 ± 0.67	16.19 ± 3.60	213.55 ± 27.59
樟子松林 Pinus sylvestris var. mongolica forest	幼龄林 Young forest	15.17 ± 2.10	1.74 ± 0.24	0.68 ± 0.18	1.07 ± 0.24	2.44 ± 0.37	21.10 ± 2.34	10.21 ± 1.51	12.67 ± 2.46	3.27 ± 0.59	47.25 ± 2.85
	中龄林 Middle-aged forest	70.11 ± 16.98	6.31 ± 0.97	1.72 ± 0.34	2.36 ± 0.57	8.25 ± 1.28	88.75 ± 17.12	4.37 ± 0.64	7.97 ± 1.10	5.16 ± 1.00	106.25 ± 19.45
	近熟林 Near mature forest	94.07 ± 13.56	8.07 ± 1.74	2.06 ± 0.38	2.74 ± 0.45	10.43 ± 2.19	117.37 ± 14.62	7.67 ± 1.27	7.78 ± 1.19	10.27 ± 1.59	143.09 ± 16.51
	成熟林 Mature forest	129.70 ± 19.99	10.58 ± 2.18	2.50 ± 0.51	3.23 ± 0.67	13.46 ± 2.26	159.47 ± 20.16	6.63 ± 1.67	6.97 ± 0.98	11.94 ± 2.45	185.01 ± 23.34
云冷杉林 Picea-Abies forest	幼龄林 Young forest	22.43 ± 1.57	1.19 ± 0.28	0.71 ± 0.24	0.86 ± 0.10	1.84 ± 0.26	27.04 ± 1.89	6.57 ± 1.66	0.41 ± 0.08	11.54 ± 2.16	45.56 ± 2.01
	中龄林 Middle-aged forest	53.31 ± 9.85	2.69 ± 0.31	2.01 ± 0.39	1.79 ± 0.38	5.18 ± 1.01	64.98 ± 10.85	5.17 ± 1.28	0.23 ± 0.03	15.57 ± 2.85	85.95 ± 12.32
	近熟林 Near mature forest	96.29 ± 10.23	4.67 ± 0.86	4.11 ± 0.85	2.94 ± 0.96	10.52 ± 2.27	118.52 ± 11.32	9.47 ± 2.94	2.54 ± 0.34	18.37 ± 2.64	148.9 ± 11.25
	成熟林 Mature forest	174.50 ± 30.42	8.15 ± 1.37	8.43 ± 0.64	4.85 ± 1.06	21.41 ± 3.48	217.34 ± 31.28	3.66 ± 0.31	2.37 ± 0.45	24.27 ± 3.88	247.64 ± 2.37
白桦林 Betula platyphylla forest	幼龄林 Young forest	22.29 ± 1.46	2.54 ± 0.89	1.15 ± 0.17	1.86 ± 0.57	3.71 ± 0.46	31.55 ± 2.00	2.45 ± 0.41	5.38 ± 0.76	1.69 ± 0.21	41.07 ± 2.37
	中龄林 Middle-aged forest	46.67 ± 7.56	4.36 ± 0.97	1.78 ± 0.29	2.83 ± 0.45	5.67 ± 1.03	61.32 ± 7.99	4.67 ± 1.21	6.87 ± 1.27	3.47 ± 0.79	76.33 ± 9.45
	近熟林 Near mature forest	91.27 ± 11.39	7.12 ± 1.46	2.64 ± 0.64	4.16 ± 0.67	8.33 ± 2.18	113.52 ± 12.56	10.11 ± 2.16	9.57 ± 1.61	9.29 ± 1.64	142.49 ± 14.00
	成熟林 Mature forest	134.09 ± 16.96	9.43 ± 2.01	3.31 ± 0.54	5.18 ± 1.56	10.39 ± 0.29	162.40 ± 17.85	12.61 ± 2.57	2.14 ± 0.53	10.57 ± 1.57	187.72 ± 19.56
蒙古栎林 Quercus mongolica forest	幼龄林 Young forest	6.82 ± 0.75	1.52 ± 0.34	0.77 ± 0.13	1.01 ± 0.18	1.35 ± 0.27	11.46 ± 1.03	8.17 ± 1.89	19.33 ± 3.59	15.78 ± 3.59	54.74 ± 11.52
	中龄林 Middle-aged forest	20.53 ± 2.14	3.75 ± 0.84	1.77 ± 0.49	2.30 ± 0.41	3.13 ± 0.26	31.48 ± 2.75	7.64 ± 1.67	0.58 ± 0.09	2.67 ± 0.44	42.37 ± 5.16
	近熟林 Near mature forest	66.63 ± 16.48	9.83 ± 1.29	4.32 ± 1.04	5.54 ± 0.86	7.71 ± 1.08	94.04 ± 17.59	2.54 ± 0.49	0.47 ± 0.06	1.33 ± 0.22	98.38 ± 18.23
	成熟林 Mature forest	100.15 ± 17.56	13.72 ± 2.18	5.89 ± 1.15	7.52 ± 1.34	10.53 ± 2.19	137.80 ± 19.75	1.37 ± 0.35	0.51 ± 0.05	1.64 ± 0.34	141.32 ± 24.56
山杨林 Populus davidiana forest	幼龄林 Young forest	18.47 ± 1.75	1.77 ± 0.19	0.78 ± 0.13	1.32 ± 0.25	2.19 ± 0.52	24.54 ± 1.86	10.21 ± 2.57	8.28 ± 2.11	1.14 ± 0.12	44.17 ± 2.31
	中龄林 Middle-aged forest	35.81 ± 5.14	3.46 ± 0.61	1.52 ± 0.27	2.16 ± 0.37	3.38 ± 0.49	46.33 ± 6.15	8.46 ± 1.84	5.37 ± 1.35	3.47 ± 0.68	63.63 ± 6.87
	近熟林 Near mature forest	63.32 ± 8.79	6.15 ± 0.91	2.69 ± 0.61	3.30 ± 0.86	4.91 ± 0.84	80.36 ± 9.42	4.96 ± 0.78	6.57 ± 1.64	4.75 ± 0.93	96.64 ± 10.34
	成熟林 Mature forest	113.41 ± 16.34	11.08 ± 1.93	4.82 ± 0.49	5.08 ± 1.24	7.20 ± 1.42	141.59 ± 17.52	9.34 ± 2.47	8.39 ± 2.02	6.34 ± 1.24	165.66 ± 19.46

林型 Forest type	龄组 Age	乔木层Tree layer						灌木层 Shrub layer	草本层 Herbaceous layer	凋落物层 Litter layer	林分碳密度 Forest stand carbon density
林型 Forest type	龄组 Age	干 Trunk	枝 Branch	叶 Leaf	皮 Bark	根 Root	小计 Total	灌木层 Shrub layer	草本层 Herbaceous layer	凋落物层 Litter layer	林分碳密度 Forest stand carbon density
红松林 Pinus koraiensis forest	幼龄林 Young forest	16.4 ± 3.59	1.3 ± 0.27	0.6 ± 0.23	0.9 ± 0.47	1.0 ± 0.45	20.2 ± 3.63	5.3 ± 2.19	1.5 ± 0.70	4.4 ± 2.07	31.4 ± 4.31
	中龄林 Middle-aged forest	52.8 ± 11.89	3.0 ± 1.11	1.5 ± 0.58	1.9 ± 0.91	2.6 ± 0.70	61.7 ± 13.94	6.2 ± 1.90	2.1 ± 0.96	4.7 ± 1.21	74.7 ± 15.75
	近熟林 Near mature forest	89.9 ± 23.48	4.2 ± 1.64	2.2 ± 1.38	2.6 ± 1.36	4.0 ± 1.27	102.9 ± 26.58	7.8 ± 2.02	2.4 ± 0.89	5.3 ± 1.75	118.4 ± 29.15
	成熟林 Mature forest	100.2 ± 20.26	4.5 ± 1.40	2.4 ± 0.80	2.8 ± 1.47	4.3 ± 1.71	114.2 ± 21.76	4.2 ± 1.71	2.6 ± 0.77	9.2 ± 2.89	130.2 ± 23.10
兴安落叶松林 Larix gmelinii forest	幼龄林 Young forest	6.7 ± 1.86	0.7 ± 0.19	0.4 ± 0.38	1.0 ± 0.35	2.1 ± 0.89	10.9 ± 2.54	9.3 ± 4.22	3.5 ± 1.49	5.1 ± 2.49	28.9 ± 2.98
	中龄林 Middle-aged forest	18.3 ± 4.67	2.0 ± 0.85	0.7 ± 0.59	1.8 ± 0.89	4.1 ± 1.30	27.0 ± 5.51	4.8 ± 1.56	5.2 ± 1.42	7.3 ± 3.48	44.3 ± 7.51
	近熟林 Near mature forest	43.4 ± 17.36	4.9 ± 1.74	1.3 ± 0.36	3.1 ± 1.30	7.4 ± 2.25	60.0 ± 18.87	2.2 ± 0.29	4.2 ± 1.77	7.8 ± 3.36	74.2 ± 20.69
	成熟林 Mature forest	76.5 ± 27.19	8.7 ± 3.89	1.8 ± 0.85	4.4 ± 1.81	10.9 ± 2.92	102.3 ± 25.97	1.2 ± 0.17	1.5 ± 0.69	8.3 ± 3.65	113.3 ± 27.86
樟子松林 Pinus sylvestris var. mongolica forest	幼龄林 Young forest	7.5 ± 2.56	0.9 ± 0.27	0.3 ± 0.20	0.5 ± 0.28	1.2 ± 0.41	10.4 ± 2.57	5.1 ± 1.53	5.8 ± 2.48	1.5 ± 0.61	22.8 ± 3.15
	中龄林 Middle-aged forest	34.5 ± 17.69	3.1 ± 1.08	0.8 ± 0.39	1.2 ± 0.60	4.1 ± 1.32	43.7 ± 17.36	2.2 ± 0.68	3.7 ± 1.12	2.4 ± 1.03	52.0 ± 20.37
	近熟林 Near mature forest	47.4 ± 14.68	4.0 ± 1.87	1.0 ± 0.41	1.3 ± 0.48	5.2 ± 2.22	58.9 ± 14.86	3.8 ± 1.29	3.6 ± 1.21	4.8 ± 1.62	71.1 ± 16.85
	成熟林 Mature forest	65.9 ± 20.45	5.2 ± 2.58	1.2 ± 0.55	1.6 ± 0.71	6.6 ± 2.29	80.5 ± 20.37	3.3 ± 1.70	3.3 ± 1.00	5.6 ± 2.48	92.6 ± 23.89
云冷杉林 Picea-Abies forest	幼龄林 Young forest	11.5 ± 1.63	0.6 ± 0.36	0.4 ± 0.28	0.4 ± 0.12	0.9 ± 0.28	13.9 ± 1.94	3.3 ± 1.69	0.2 ± 0.12	5.8 ± 2.18	23.1 ± 2.67
	中龄林 Middle-aged forest	27.6 ± 9.99	1.4 ± 0.36	1.0 ± 0.42	0.9 ± 0.41	2.7 ± 1.06	33.6 ± 10.96	2.6 ± 1.31	0.1 ± 0.06	7.9 ± 2.91	44.1 ± 12.86
	近熟林 Near mature forest	50.5 ± 10.54	2.4 ± 0.91	2.1 ± 0.88	1.5 ± 0.98	5.5 ± 2.32	62.1 ± 11.51	4.9 ± 2.97	1.2 ± 0.37	9.5 ± 2.67	77.6 ± 11.97
	成熟林 Mature forest	92.8 ± 30.87	4.2 ± 1.42	4.3 ± 0.67	2.5 ± 1.09	11.3 ± 3.52	115.1 ± 31.49	1.9 ± 0.36	1.1 ± 0.48	12.3 ± 3.98	130.3 ± 33.56
白桦林 Betula platyphylla forest	幼龄林 Young forest	10.4 ± 1.58	1.2 ± 0.92	0.5 ± 0.20	0.8 ± 0.60	1.7 ± 0.48	14.5 ± 2.33	1.1 ± 0.44	2.4 ± 0.79	0.8 ± 0.23	18.8 ± 2.88
	中龄林 Middle-aged forest	21.9 ± 7.88	2.0 ± 0.99	0.8 ± 0.32	1.3 ± 0.47	2.6 ± 1.05	28.5 ± 8.49	2.2 ± 1.27	3.1 ± 1.30	1.6 ± 0.83	35.3 ± 10.47
	近熟林 Near mature forest	43.2 ± 11.54	3.3 ± 1.50	1.2 ± 0.66	1.9 ± 0.69	3.8 ± 2.19	53.3 ± 12.87	4.7 ± 2.18	4.3 ± 1.64	4.3 ± 1.69	66.6 ± 14.85
	成熟林 Mature forest	63.8 ± 17.25	4.4 ± 2.06	1.5 ± 0.57	2.3 ± 1.58	4.8 ± 0.31	76.8 ± 18.16	6.0 ± 2.59	1.0 ± 0.58	4.9 ± 1.59	88.5 ± 20.37
蒙古栎林 Quercus mongolica forest	幼龄林 Young forest	3.2 ± 0.84	0.7 ± 0.40	0.4 ± 0.16	0.5 ± 0.20	0.6 ± 0.30	5.4 ± 1.34	3.9 ± 1.92	8.7 ± 3.62	7.0 ± 3.62	25.0 ± 12.69
	中龄林 Middle-aged forest	9.8 ± 2.19	1.8 ± 0.88	0.8 ± 0.52	1.1 ± 0.44	1.5 ± 0.27	14.9 ± 2.87	3.6 ± 1.69	0.3 ± 0.10	1.2 ± 0.47	20.0 ± 6.16
	近熟林 Near mature forest	32.5 ± 16.87	4.7 ± 1.31	2.0 ± 1.07	2.6 ± 0.88	3.6 ± 1.11	45.4 ± 17.76	1.2 ± 0.52	0.2 ± 0.07	0.6 ± 0.25	47.5 ± 19.42
	成熟林 Mature forest	49.4 ± 17.94	6.5 ± 2.22	2.8 ± 1.18	3.6 ± 1.36	5.0 ± 2.24	67.2 ± 19.86	0.7 ± 0.38	0.2 ± 0.07	0.8 ± 0.38	68.9 ± 25.26
山杨林 Populus davidiana forest	幼龄林 Young forest	8.3 ± 1.76	0.8 ± 0.20	0.3 ± 0.16	0.6 ± 0.26	1.0 ± 0.55	11.0 ± 1.99	4.6 ± 2.59	3.7 ± 2.14	0.5 ± 0.15	19.8 ± 2.58
	中龄林 Middle-aged forest	16.2 ± 5.19	1.6 ± 0.67	0.7 ± 0.30	1.0 ± 0.38	1.6 ± 0.52	20.9 ± 6.58	3.9 ± 1.87	2.4 ± 1.37	1.5 ± 0.70	28.7 ± 7.19
	近熟林 Near mature forest	28.7 ± 9.42	2.8 ± 0.94	1.2 ± 0.63	1.5 ± 0.91	2.2 ± 0.88	36.3 ± 9.64	2.3 ± 0.82	3.0 ± 1.68	2.1 ± 0.95	43.7 ± 11.75
	成熟林Mature forest	52.8 ± 16.83	5.1 ± 1.98	2.2 ± 0.51	2.3 ± 1.25	3.3 ± 1.46	65.7 ± 17.87	4.3 ± 2.51	3.8 ± 2.05	2.9 ± 1.26	76.6 ± 20.45

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